Professional Documents
Culture Documents
P. kickapooensis
O. c/arki
.. ------- --------- 11:01le'a- - - --
P. k;ckapooensis w
I:
C ex gr. triJamra P. rsussi G. obesa-G. levis
Cassidella ex gr.
P arena!a P. plummerse
G.obess viscidus
-------
;- - - - - - --
iOBiter/er - - - -
------ --------
---------
.g
A. impexu5
O. ve/ascoensis A. paterslla
..
i
O. vslascoensis
..
G.l8vis-
A. imp8XUS
Discammina sp .A
10
1-
------- ---------
8X-g,:-b8aumont;anus
...J Discammina sp. A
A. patereJJa
Discammina sp .A -
C. ex Qr.
G. nonionoides Gabonita levis
beaumontJana- P. ex gr. prolixa
------- -------- --------
w
G, nonionoides
I A...;. cl _
P. ex gr. fang I:
NOdOSaria ex. gr. Cibicides sp.A
.!II _ ____
I: obscura-
. supracretsces .!!
CiblcK1es sp, A
_ _
-------
--------- G. corrects
L. tormarpensis
G.lorneiana
I:
G. sandidgei
G. nacatochensis
R . ex gr. szajnochae
..
w G. subconica 10 ------ --------
----------
G. loetterlei
...J
N. subcrelacea- N. subcretacea
S.
H. ex gr. subsphaerica
S. et. crassicosla S. et. crassicostata
R. ex gr, szajnochae
------ --------
---------
G. berthefini-
0. schloenbachi
S. bramJettei lJIum.merae-
G. berthelini-
T!!aE.e!i. _____
---------
r-- rsussl p1exus-
plummerae...
------- G. cf.gradata
'!.u:,sij''!.x,:s _ _ _
G. cf. gradata
ill"" ".
o1l
------
---------
Glaphyrammina sp. A .., E. carpenteri
O. schloenbachi-
T. ex gr. excavata
PIS ex gr. dividens
ex gr bulleta
PIS. ex gr. divid&ns
>- O. sch/oenbachi
iii
G. spissocostara
w
R. ex gr. globulosa
------ --------
---------
w
8.
L revoIU'B-
rr ;p-A
E'J.arcrcw:Z-
G. filiformls
G, spinea
L. revoluta E. carpenteri
E. aracajuensis
G./Hiformis
!
G. ex gr. beccariiformis
------
6! _
---------
I: G. correcta
su';,:rr.:;a-
L. ex gr. nodosa
.!II G. sandldgei
L. ex gr. sUbangulata
j Gavelinopsis(?) sp. B L. ex gr. nodosa
B. nonioninoides
I: L. revoluta I:
--A.cT-- --------
.3
P ex gr. capitosa .. .!II
A. cf. coprofithiformis
,-
------ loo- --------
j
coprolithiformis-
8. hedbe'9i
GaveJlne7Ja-sp B - -
8 cl. hedber9i-
H.lueckel
H.luecksl
>-
Gavel/inella sp. A-
L formarpensls
-"LCi';---
L. nodosaria
1.i Valvulineria sp. A T. ex r SUbCOnlC8 L. ciryi
w
sp B
L. nodosaria-
__ J,f-
M. ex gr. aequivocs
-------
Fig. 7 - FAD and LAD ofbiostratigraphlcally important benthlc Foraminiferlda (age lengths not to scale).
Owing to the limited vertical extent of most outcrop sections (nonnally only a few metres) in relation
to the considerable total thickness of the carbonate sequence (well over 500 m), no significant faunal shifts
were nonnally observed during macrofossil sampling of each locality. Bed-by-bed sampling was therefore
not consideredpractical; however, as a result of this broad collecting procedure some samples may represent
mixed assemblages.
Macrofossil sampling in Sergipe is hampered by lackof continuous outcrop. Nonnally small quarries
and road cuttings expose only up to a few metres of section, and initial collecting procedures yielded only a
small number of specimens. The few existing larger quarries have in many cases yielded sizeable collections
252
BENTHIC FORAMINIFERA
-------
---------
ValvulineriB
G-: berthe7inr: - - -
..
sp.
plummeras-
Ages Zones
First app. Last app. 10
plummerae-- reU$S; plexus
datums datums
...J
c:
_ _ _
---------
1--:-
.!II
G./evis
c:
B. ex gr. colonensis
..,
e G.f8vis-
N. ex gr.obscura
N. aspers
I.....!...
N. ex gr. obscura Gaboni!a cf. paNa
O. clarki-
N. canadenss
P. kickapooensis
O. clarki
.. ------- --------- 11:01le'o- - - --
P. k;ckapooensis w
c:
C ex gr. tri/atera P. fsussi G. obesa-G. levis
Cassidella ex gr.
P arena!a P. plummerse
G. abess viscidus
-------
;- - - - - - --
ioeffe,l"r - - - -
------ --------
---------
.g
A. impexus
O. ve/ascoensis A. paterslla
..
i
O. vslascoensis
..
G.l8vis-
A. impexus
Discammina sp .A
10
1-
------- ---------
8X-g,:-b80umont;onus
...J Discammina sp, A A. patereJJa
Discammina sp .A -
C. ex Qf.
G. nonionoides Gabonita levis
beaumontJana- P. ex gr. prolixa
------- -------- --------
w
G, nonionoides
I A...;. c!: _
P. ex gr. fang c:
NOdOSaria ex. gr. Cibicides sp.A
.!II _ ____
c: obscura-
E. supracretsces .!!
CiblcK1es sp, A
_ _
-------
--------- G. corrects
L. tormarpensis
G./orneiana
c:
G. sandidge;
G. nacatochensis
R . ex gr. szajnochae
..
w G. subconica 10 ------ --------
----------
G. /oetterlei
...J
N. subcrelacea- N. subcretacea
S.
H. ex gr. subsphaerica
S. cf. crassicosla S. cf. crassicostata
R. ex gr, szajnochae
------ --------
---------
G. berthefini-
0. schloenbachi
S. bramJettei lJIum.merae-
G. berthelini-
T!!aE.e!i. _____
---------
r-- rsuss/ p1exus-
p/ummerae...
------- G. cf.gradata
'!.u:,sij''!.x,:s _ _ _
G. ct. gradata
ill"" ".
o1l
------
---------
Glaphyrammina sp. A .., E. carpenteri
O. schloenbachi-
T. ex gr. excavata
PIS ex gr. dividens
ex gr bulleto
PIS. ex gr. divid&ns
>- O. sch/oenbachi
iii
G. spissocostara
w
R. ex gr. globulosa
------ --------
---------
w
8.
L revoIU'B-
rr ;p-A
E'J.arcrcw:Z-
G. fili/ormls
G, spinea
L. rBvo/uta E. carpenter;
E. aracajuensis
G.lili/ormis
!
G. ex gr. beccariiformis
------
6! _
---------
c: G. correcta
su';,:rr.:;a-
L. ex gr. nodosa
.!II G. sandldgei
L. ex gr. sUbangulata
j Gavelinopsis(?) sp. B L. ex gr. nodosa
B. nonioninoides
c: L. revo/uta c:
--A.cT-- --------
.3
P ex gr. capitosa .. .!II
A. cf. coprofithiformis
,-
------ fo. --------
j
coprolithiformis-
8. hedbe'9i
GoveJlne7Jo-sp B - -
8 cf. hedber9i-
H. luecke,
H.luecks/
>-
Gavellinel/a sp. A-
L rormarpens/s
-"LCi';---
L. nodosaria
1.i Valvulineria sp. A T. ex r SUbCOnlC8 L. ciryi
w
sp B
L. nodosaria-
__ J,f-
M. ex gr. aequivocs
-------
Fig. 7 - FAD and LAD ofbiostratigraphlcally important benthic Foraminiferida (age lengths not to scale).
Owing to the limited vertical extent of most outcrop sections (nonnally only a few metres) in relation
to the considerable total thickness of the carbonate sequence (well over 500 m), no significant faunal shifts
were nonnally observed during macrofossil sampling of each locality. Bed-by-bed sampling was therefore
not consideredpractical; however, as a result of this broad collecting procedure some samples may represent
mixed assemblages.
Macrofossil sampling in Sergipe is hampered by lackof continuous outcrop. Nonnally small quarries
and road cuttings expose only up to a few metres of section, and initial collecting procedures yielded only a
small number of specimens. The few existing larger quarries have in many cases yielded sizeable collections
252
BENTHIC FORAMINIFERA
-------
---------
ValvulineriB
G-: berthe7inr: - - -
..
sp.
plummeras-
Ages Zones
First app. Last app. 10
plummerae-- reU$S; plexus
datums datums
...J
I:
_ _ _
---------
1--:-
.!II
G./evis
I:
B. ex gr. colonensis
..,
e G.f8vis-
N. ex 9r. obscura
N. aspsrs
I.....!...
N. ex gr. obscura Gaboni!a cf. paNa
O. clarki-
N. canadenss
P. kickapooensis
O. c/arki
.. ------- --------- 11:01le'a- - - --
P. k;ckapooensis w
I:
C ex gr. triJamra P. rsussi G. obesa-G. levis
Cassidella ex gr.
P arena!a P. plummerse
G.obess viscidus
-------
;- - - - - - --
iOBiter/er - - - -
------ --------
---------
.g
A. impexu5
O. ve/ascoensis A. paterslla
..
i
O. vslascoensis
..
G.l8vis-
A. imp8XUS
Discammina sp .A
10
1-
------- ---------
8X-g,:-b8aumont;anus
...J Discammina sp. A
A. patereJJa
Discammina sp .A -
C. ex Qr.
G. nonionoides Gabonita levis
beaumontJana- P. ex gr. prolixa
------- -------- --------
w
G, nonionoides
I A...;. cl _
P. ex gr. fang I:
NOdOSaria ex. gr. Cibicides sp.A
.!II _ ____
I: obscura-
. supracretsces .!!
CiblcK1es sp, A
_ _
-------
--------- G. corrects
L. tormarpensis
G.lorneiana
I:
G. sandidgei
G. nacatochensis
R . ex gr. szajnochae
..
w G. subconica 10 ------ --------
----------
G. loetterlei
...J
N. subcrelacea- N. subcretacea
S.
H. ex gr. subsphaerica
S. et. crassicosla S. et. crassicostata
R. ex gr, szajnochae
------ --------
---------
G. berthefini-
0. schloenbachi
S. bramJettei lJIum.merae-
G. berthelini-
T!!aE.e!i. _____
---------
r-- rsussl p1exus-
plummerae...
------- G. cf.gradata
'!.u:,sij''!.x,:s _ _ _
G. cf. gradata
ill"" ".
o1l
------
---------
Glaphyrammina sp. A .., E. carpenteri
O. schloenbachi-
T. ex gr. excavata
PIS ex gr. dividens
ex gr bulleta
PIS. ex gr. divid&ns
>- O. sch/oenbachi
iii
G. spissocostara
w
R. ex gr. globulosa
------ --------
---------
w
8.
L revoIU'B-
rr ;p-A
E'J.arcrcw:Z-
G. filiformls
G, spinea
L. revoluta E. carpenteri
E. aracajuensis
G./Hiformis
!
G. ex gr. beccariiformis
------
6! _
---------
I: G. correcta
su';,:rr.:;a-
L. ex gr. nodosa
.!II G. sandldgei
L. ex gr. sUbangulata
j Gavelinopsis(?) sp. B L. ex gr. nodosa
B. nonioninoides
I: L. revoluta I:
--A.cT-- --------
.3
P ex gr. capitosa .. .!II
A. cf. coprofithiformis
,-
------ loo- --------
j
coprolithiformis-
8. hedbe'9i
GaveJlne7Ja-sp B - -
8 cl. hedber9i-
H.lueckel
H.luecksl
>-
Gavel/inella sp. A-
L formarpensls
-"LCi';---
L. nodosaria
1.i Valvulineria sp. A T. ex r SUbCOnlC8 L. ciryi
w
sp B
L. nodosaria-
__ J,f-
M. ex gr. aequivocs
-------
Fig. 7 - FAD and LAD ofbiostratigraphlcally important benthlc Foraminiferlda (age lengths not to scale).
Owing to the limited vertical extent of most outcrop sections (nonnally only a few metres) in relation
to the considerable total thickness of the carbonate sequence (well over 500 m), no significant faunal shifts
were nonnally observed during macrofossil sampling of each locality. Bed-by-bed sampling was therefore
not consideredpractical; however, as a result of this broad collecting procedure some samples may represent
mixed assemblages.
Macrofossil sampling in Sergipe is hampered by lackof continuous outcrop. Nonnally small quarries
and road cuttings expose only up to a few metres of section, and initial collecting procedures yielded only a
small number of specimens. The few existing larger quarries have in many cases yielded sizeable collections
252
of biostratigraphically significant macrofossils ; however, because the material was collected chiefly by
quarry workers, the relative position in sequence of most of the specimens is unknown. Position in sequence
of isolated localities was inferred mainly on the basis of regional dip and altitude of the beds exposed. Thus,
in contrast to microbiostratigraphical procedures, the true first appearances and the true vertical distribution
of the diagnostic macrofossil taxa have not been observed.
For the definition of the base ofeach ammonite zone, the first appearance of a well-known, abundant,
wide-spread, and easily identifiable taxon was chosen. In addition to the defining taxon, a number of other
taxa characterize the zone and thus aid in its identification.
Reference sections and profiles for the biownes will be given in a forthcoming publication (Koutsou-
kos & Bengtson, in preparation)..
Legend - The abbreviations given after the diagnostic taxa listed represent local biotic events and
occurrences, recorded at outcrop and in well-sections in the Sergipe Basin.
(FAD) =First Appearance Datum
(FAWZ) = First Appearance Within Zone
(LAD) =Last Appearance Datum
(LAWZ) =Last Appearance Within Zone
(TR) = Total Range
(AC) = ACme (abundance peak)
(OWZ) =Occurrence Within Zone
(OWZPR) = Occurrence Within Zone, Probably Restricted to zone
PLANKTONIC FORAMINIFERIDA
The succession of planktonic foraminiferids is based in part on the works of Masters (1977),
Robaszynski & Caron (1979), Robaszynski et al. (1984) and Caron (1985).
Globigerinelloides barri-Hedbergella (H.) gorbachikae Zone: Hedbergella (H.) trocoidea (Gandol-
fi, 1942) (FAWZ), Hedbergella (H.) gorbachikae Longoria, 1974 (FAWZ), Globigerinelloides barri (Bolli,
Loeblich & Tappan, 1957) (OWZ). Assigned age: late Aptian.
Globigerinelloides ex gr. maridalensis-Hedbergella (H.) similis Zone: Ticinella spp. (FAD), Globi-
gerinelloids barri (Bolli, Loeblich & Tappan, 1957) (LAD), Globigerinelloidsferreolensis (Moullade, 1961)
(LAD), Globigerinelloids ex gr. maridalensis (Bolli, 1959) (LAD), Hedbergella (H.) similis Longoria, 1974
(LAD), Ticinella bejaouaensis Sigal, 1966 (FAD, AC; "1st abundance peak"). Assigned age: late Aptian.
Globigerinelloides cushmani-Ticinella bejaouaensis Zone: Globigerinelloids cushmani (Tappan,
1943) (FAD), Ticinella bejaouaensis Sigal, 1966(AC; "2ndabundancepeak"). Assignedage : earliest Albian.
Ticinella bejaouaensis Zone: Category: Acme-wne (sensu Hedberg, 1976, p. 59-60). Hedbergella
(H.) subcretacea (Tappan, 1943) (FAD), Ticinella bejaouaensis Sigal, 1966 (AC ; "3rd abundance peak").
Assigned age: early Albian.
Tiein'ella ex gr. primula-Tieinella bejaouaensis Zone: Hedbergella (H.) ex gr. simplex (Morrow,
1934) (FAD), Globigerinelloides bentonensis (Morrow, 1934) (FAD), Ticinella ex gr.primulaLuterbacher,
1963 (FAD), Globigerinelloides cushmani (Tappan, 1943) (LAD), Ticinella bejaouaensis Sigal, 1966
(LAD). Assigned age: mid-Albian.
Bi/ieinella breggiensis-Tieinella ex gr. primula Zone: Biticinella breggiensis (Gandolfi, 1942)
(FAD), Ticinella raynaudi Sigal, 1966 (FAD), Ticinella madecassiana Sigal, 1966 (TR?), large-sized
Ticinellinae (AC ; 4th abundance peak; in the upper limit). Assigned age: mid to late Albian.
Globigerinelloides texomaensis-Bitieinella breggiensis Zone: Globigerinelloides texomaensisMi-
chael, 1972 (FAD), Praeglobotruncana delrioensis (plummer, 1931) (FAD), Heterohelix spp. (FAD),
Biticinella breggiensis (Gandolfi, 1942) (LAD), Ticinella ex gr. primulaLuterbacher, 1963 (LAD). Assigned
age: late Albian.
Hedbergella (H.) gorbachikae-Ticinella raynaudi Zone: Guembelitria cenomana (Keller, 1935)
(FAD), Globigerinelloides texomaensis Michael, 1972 (LAD), Hedbergella (H.) gorbachikae Longoria,
1974 (LAD), Ticinella spp. (T. raynaudi Sigal, 1966, T. roberti (Gandolfi, 1942); (LAD). Assignedage : latest
Albian.
253
of biostratigraphically significant macrofossils; however, because the material was collected chiefly by
quarry workers, the relative position in sequence of most of the specimens is unknown. Position in sequence
of isolated localities was inferred mainly on the basis of regional dip and altitude of the beds exposed. Thus,
in contrast to microbiostratigraphical procedures, the true first appearances and the true vertical distribution
of the diagnostic macrofossil taxa have not been observed.
For the definition of the base ofeach ammonite zone, the first appearance of a well-known, abundant,
wide-spread, and easily identifiable taxon was chosen. In addition to the defining taxon, a number of other
taxa characterize the zone and thus aid in its identification.
Reference sections and profiles for the biownes will be given in a forthcoming publication (Koutsou-
kos & Bengtson, in preparation)..
Legend - The abbreviations given after the diagnostic taxa listed represent local biotic events and
occurrences, recorded at outcrop and in well-sections in the Sergipe Basin.
(FAD) = First Appearance Datum
(FAWZ) = First Appearance Within Zone
(LAD) = Last Appearance Datum
(LAWZ) = Last Appearance Within Zone
(TR) = Total Range
(AC) = ACme (abundance peak)
(OWZ) = Occurrence Within Zone
(OWZPR) = Occurrence Within Zone, Probably Restricted to zone
PLANKTONIC FORAMINIFERIDA
The succession of planktonic foraminiferids is based in part on the works of Masters (1977),
Robaszynski & Caron (1979), Robaszynski et al. (1984) and Caron (1985).
Globigerinelloides barri-Hedbergella (H.) gorbachikae Zone: Hedbergella (H.) trocoidea (Gandol-
fi, 1942) (FAWZ), Hedbergella (H.) gorbachikae Longoria, 1974 (FAWZ), Globigerinelloides barri (Bolli,
Loeblich & Tappan, 1957) (OWZ). Assigned age: late Aptian.
Globigerinelloides ex gr. maridalensis-Hedbergella (H.) similis Zone: Ticinella spp. (FAD), Globi-
gerinelloids barri (Bolli, Loeblich & Tappan, 1957) (LAD), Globigerinelloidsferreolensis (Moullade, 1961)
(LAD), Globigerinelloids ex gr. maridalensis (Bolli, 1959) (LAD), Hedbergella (H.) similis Longoria, 1974
(LAD), Ticinella bejaouaensis Sigal, 1966 (FAD, AC; "1st abundance peak"). Assigned age: late Aptian.
Globigerinelloides cushmani-Ticinella bejaouaensis Zone: Globigerinelloids cushmani (Tappan,
1943) (FAD), Ticinella bejaouaensis Sigal, 1966(AC; "2ndabundancepeak"). Assignedage : earliest Albian.
Ticinella bejaouaensis Zone: Category: Acme-wne (sensu Hedberg, 1976, p. 59-60). Hedbergella
(H.) subcretacea (Tappan, 1943) (FAD), Ticinella bejaouaensis Sigal, 1966 (AC ; "3rd abundance peak").
Assigned age: early Albian.
Ticin'ella ex gr. primula-Ticinella bejaouaensis Zone: Hedbergella (H.) ex gr. simplex (Morrow,
1934) (FAD), Globigerinelloides bentonensis (Morrow, 1934) (FAD), Ticinella ex gr.primulaLuterbacher,
1963 (FAD), Globigerinelloides cushmani (Tappan, 1943) (LAD), Ticinella bejaouaensis Sigal, 1966
(LAD). Assigned age: mid-Albian.
Biticinella breggiensis-Ticinella ex gr. primula Zone: Biticinella breggiensis (Gandolfi, 1942)
(FAD), Ticinella raynaudi Sigal, 1966 (FAD), Ticinella madecassiana Sigal, 1966 (TR?), large-sized
Ticinellinae (AC ; 4th abundance peak; in the upper limit). Assigned age: mid to late Albian.
Globigerinelloides texomaensis-Biticinella breggiensis Zone: Globigerinelloides texomaensisMi-
chael, 1972 (FAD), Praeglobotruncana delrioensis (plummer, 1931) (FAD), Heterohelix spp. (FAD),
Biticinella breggiensis (Gandolfi, 1942) (LAD), Ticinella ex gr. primulaLuterbacher, 1963 (LAD). Assigned
age: late Albian.
Hedbergella (H.) gorbachikae-Ticinella raynaudi Zone: Guembelitria cenomana (Keller, 1935)
(FAD), Globigerinelloides texomaensis Michael, 1972 (LAD), Hedbergella (H.) gorbachikae Longoria,
1974 (LAD), Ticinella spp. (T. raynaudi Sigal, 1966, T. roberti (Gandolfi, 1942); (LAD). Assignedage : latest
Albian.
253
of biostratigraphically significant macrofossils ; however, because the material was collected chiefly by
quarry workers, the relative position in sequence of most of the specimens is unknown. Position in sequence
of isolated localities was inferred mainly on the basis of regional dip and altitude of the beds exposed. Thus,
in contrast to microbiostratigraphical procedures, the true first appearances and the true vertical distribution
of the diagnostic macrofossil taxa have not been observed.
For the definition of the base ofeach ammonite zone, the first appearance of a well-known, abundant,
wide-spread, and easily identifiable taxon was chosen. In addition to the defining taxon, a number of other
taxa characterize the zone and thus aid in its identification.
Reference sections and profiles for the biownes will be given in a forthcoming publication (Koutsou-
kos & Bengtson, in preparation)..
Legend - The abbreviations given after the diagnostic taxa listed represent local biotic events and
occurrences, recorded at outcrop and in well-sections in the Sergipe Basin.
(FAD) = First Appearance Datum
(FAWZ) = First Appearance Within Zone
(LAD) = Last Appearance Datum
(LAWZ) = Last Appearance Within Zone
(TR) = Total Range
(AC) = ACme (abundance peak)
(OWZ) = Occurrence Within Zone
(OWZPR) = Occurrence Within Zone, Probably Restricted to zone
PLANKTONIC FORAMINIFERIDA
The succession of planktonic foraminiferids is based in part on the works of Masters (1977),
Robaszynski & Caron (1979), Robaszynski et al. (1984) and Caron (1985).
Globigerinelloides barri-Hedbergella (H.) gorbachikae Zone: Hedbergella (H.) trocoidea (Gandol-
fi, 1942) (FAWZ), Hedbergella (H.) gorbachikae Longoria, 1974 (FAWZ), Globigerinelloides barri (Bolli,
Loeblich & Tappan, 1957) (OWZ). Assigned age: late Aptian.
Globigerinelloides ex gr. maridalensis-Hedbergella (H.) similis Zone: Ticinella spp. (FAD), Globi-
gerinelloids barri (Bolli, Loeblich & Tappan, 1957) (LAD), Globigerinelloidsferreolensis (Moullade, 1961)
(LAD), Globigerinelloids ex gr. maridalensis (Bolli, 1959) (LAD), Hedbergella (H.) similis Longoria, 1974
(LAD), Ticinella bejaouaensis Sigal, 1966 (FAD, AC; "1st abundance peak"). Assigned age: late Aptian.
Globigerinelloides cushmani-Ticinella bejaouaensis Zone: Globigerinelloids cushmani (Tappan,
1943) (FAD), Ticinella bejaouaensis Sigal, 1966(AC; "2ndabundancepeak"). Assignedage : earliest Albian.
Ticinella bejaouaensis Zone: Category: Acme-wne (sensu Hedberg, 1976, p. 59-60). Hedbergella
(H.) subcretacea (Tappan, 1943) (FAD), Ticinella bejaouaensis Sigal, 1966 (AC ; "3rd abundance peak").
Assigned age: early Albian.
Tiein'ella ex gr. primula-Tieinella bejaouaensis Zone: Hedbergella (H.) ex gr. simplex (Morrow,
1934) (FAD), Globigerinelloides bentonensis (Morrow, 1934) (FAD), Ticinella ex gr.primulaLuterbacher,
1963 (FAD), Globigerinelloides cushmani (Tappan, 1943) (LAD), Ticinella bejaouaensis Sigal, 1966
(LAD). Assigned age: mid-Albian.
Bi/ieinella breggiensis-Tieinella ex gr. primula Zone: Biticinella breggiensis (Gandolfi, 1942)
(FAD), Ticinella raynaudi Sigal, 1966 (FAD), Ticinella madecassiana Sigal, 1966 (TR?), large-sized
Ticinellinae (AC ; 4th abundance peak; in the upper limit). Assigned age: mid to late Albian.
Globigerinelloides texomaensis-Bitieinella breggiensis Zone: Globigerinelloides texomaensisMi-
chael, 1972 (FAD), Praeglobotruncana delrioensis (plummer, 1931) (FAD), Heterohelix spp. (FAD),
Biticinella breggiensis (Gandolfi, 1942) (LAD), Ticinella ex gr. primulaLuterbacher, 1963 (LAD). Assigned
age: late Albian.
Hedbergella (H.) gorbachikae-Ticinella raynaudi Zone: Guembelitria cenomana (Keller, 1935)
(FAD), Globigerinelloides texomaensis Michael, 1972 (LAD), Hedbergella (H.) gorbachikae Longoria,
1974 (LAD), Ticinella spp. (T. raynaudi Sigal, 1966, T. roberti (Gandolfi, 1942); (LAD). Assignedage : latest
Albian.
253
Rotaliporabrol%eni Zone: Rotaliporabrotzeni (Sigal. 1948) (FAD/LAD; lR). Assignedage: earliest
Cenomanian.
Praeglobotruncana delrioensis-Rotalipora appenninica Zone :Hedbergella (Favusella) washiten-
sis (Carsey. 1926) (LAD). Praeglobotruncana delrioensis (plummer. 1931) (LAD). Rotalipora appennini-
ca (Reoz. 1936) (LAD). Assigned age: early to mid-Cenomanian.
Hedbergella (Whiteinelltl) baltica-Hedbergella (W.) brittonensis Zone: Hedbergella (Whiteinella)
baltica Douglas &Rankin. 1969 (FAD). Hedbergella (Whiteinella) brittonensis Loeblich &Tappan. 1961
(FAD). Assigned age: mid to late Cenomanian.
Hedbergelltl (W.)aprka-Globigerinelloides bentonensis Zone: Hedbergella (Whiteinella) aprica
(Loeblich & Tappan. 1961) (FAD), Globigerinelloides bentonensis (Morrow. 1934) (LAD). Assigned age:
late Cenomanian.
Hedbergella (W.) archaeocretacea-Heterohelix reussi Zone: Hedbergella (Whiteinella) archaeo-
eretacea Pessagno. 1967 (FAD). Heterohelix reussi (Cushman. 1938) (FAWZ). Assigned age: latest Ceno-
manian to earliest Turonian.
Hedbergelltl (W.) aprka-Hedbergella (W.) baUica Z o n ~ : Marginotruncana undulata (Lehmann.
1963) (FAD). Hedbergella (Whiteinella) aprica (Loeblich & Tappan. 1961) (LAD). Hedbergella (Whitei-
nella) baltica Douglas & Rankin,l969 (LAD), Heterohelix moremani (Cushman. 1938) (LAWZ). Assigned
age: early to late Turonian.
Dicarinella primitiva Zone: Dicarinella primiliva (Dalbiez. 1955) (FAD). Guembelilria cenomana
(Keller. 1935) (LAD). Assigned age: latest Turonian.
Archaeoglobigerina cretacea-Dicarinellaprimitiva Zone: Archaeoglobigerina spp. (A. blowi Pes-
sagno. 1967, A. erelacea (d'Orbigny. 1840) ; (FAD). Dicarinella primiliva (Dalbiez. 1955) (LAD). Hedber-
gella (H.) ex gr. simplex (Morrow. 1934) (LAD). Assigned age: early to mid-Coniacian.
Dicarinelltl concavata-Marginotruncana sinuosa Zone: Marginotruncana sinuosa Porthault. 1970
(FAD). Dicarinella concavala (Brotzen. 1934) (FAD/LAD ; lR?). Hedbergella (H.) flandrini Porthault.
1970 (LAD), Marginotruncana renzi (Gandolfi. 1942) (LAWZ; late Coniacian).Marginotruncana undulata
(Lehmann. 1963) (LAWZ ; late Coniacian). Conlusolruncana ex gr.fornicata (plummer. 1931) (FAWZ;
Santonian). Ventilabrella auslinana Cushman. 1938 (FAWZ; Santonian). Assigned age: late Coniacian to
early-late Santonian.
Dicarinella asymetrica Zone: Dicarinella asymelrica (Sigal. 1952) (FAD/LAD; lR?), Hedbergel-
la (H.) ex gr. delrioensis (Carsey. 1926) (LAD). Assigned age: late Santonian.
Contusotruncana patellijormis-Globotruncanita elevata-stuartijormis plexus Zone :
Contusotruncanapatelliformis (Gandolfi, 1955) (FAD). Globotruncanitaelevata (Brotzen. 1934) - G. stuar-
tiformis (Dalbiez, 1955) plexus (FAD/LAD; lR?). G/obotruncanella havanensis (Voorwijk. 1937) (FAD).
G/obotruncanita stuartiformis S.s. (FAD), Heterohe/ix reussi (Cushman) (LAWZ). Assigned age: early
Campanian.
Globotruncana orientaUs-Globotruncana ventricosa Zone: Globotruncana linneiana (d'Orbigny.
1839) (FAD). G. bu/loides Vogler, 1941 (FAD). G. orienta/is El Naggar. 1966 (FAD). G. ventricosa White.
1928 (FAD). Globotruncanita subspinosa (pessagno. 1960) (FAD). P/anog/obu/ina varians (Rzehak. 1895)
(FAD). Pseudotextu/aria e/egans (Rzehak, 1891) (FAD). Rugog/obigerina ex gr. rugosa (plummer. 1926)
(FAD). G/obotruncanita stuarti (de Lapparent. 1918) (FAD). Assigned age: early-late Campanian.
Contusotruncana ex gr. jornicata-Globotruncana linneiana Zone: Pseudotextularia browni
Masters. 1976 (FAD). Pseudoguembelina palpebra Bronnimann & Brown, 1953 (FAD), Contusotruncana
ex gc. fornicata (plummer. 1931) (LAD), G/obotruncana bu/loides Vogler, 1941 (LAD). G. /inneiana
(d'Orbigny. 1839) (LAD). Globotruncanita subspinosa (pessagno. 1960) (LAD). Gansserina gansseri
(Bolli. 1951) (FAWZ). Globotruncana aegyptiaca Nakkady, 1950 (FAWZ), G/obotruncanella peta/oidea
(Gandolfi. 1955) (FAWZ). Assigned age: early Maastrichtian.
Gansserinagansseri-Globotruncanitastuartiformis Zone: Contusotruncanapatelliformis (Gandol-
fi, 1955) (LAD). Globotruncanita pettersi (Gandolfi. 1955) (FAD), Guembe/itria eretacea Cushman. 1933
(FAD). Gansserina gansseri (Bolli. 1951) (LAD). G/obotruncana rosetta (Carsey. 1926) (LAD). G.
ventricosaWhite. 1928(LAD). G/obotruncanita stuartiformis (Dalbiez. 1955) S.s. (LAD). Assignedage: late
Maastrichtian.
Contusotruncana contusa-Globotruncana aegyptiaca Zone: Contusotruncana contusa (Cushman,
1926) (LFAD/LLAD ; lR). Globotruncana aegyptiaca Nakkady. 1950 (LAD). P/anoglobu/ina varians
254
Rotaliporabrol%eni Zone: Rotaliporabrotzeni (Sigal. 1948) (FAD/LAD; lR). Assignedage: earliest
Cenomanian.
Praeglobotruncana delrioensis-Rotalipora appenninica Zone :Hedbergella (Favusella) washiten-
sis (Carsey. 1926) (LAD). Praeglobotruncana delrioensis (plummer. 1931) (LAD). Rotalipora appennini-
ca (Reoz. 1936) (LAD). Assigned age: early to mid-Cenomanian.
Hedbergella (Whiteinelltl) baltica-Hedbergella (W.) brittonensis Zone: Hedbergella (Whiteinella)
baltica Douglas & Rankin. 1969 (FAD). Hedbergella (Whiteinella) brittonensis Loeblich & Tappan. 1961
(FAD). Assigned age: mid to late Cenomanian.
Hedbergelltl (W.)aprka-Globigerinelloides bentonensis Zone: Hedbergella (Whiteinella) aprica
(Loeblich & Tappan. 1961) (FAD), Globigerinelloides bentonensis (Morrow. 1934) (LAD). Assigned age:
late Cenomanian.
Hedbergella (W.) archaeocretacea-Heterohelix reussi Zone: Hedbergella (Whiteinella) archaeo-
cretacea Pessagno. 1967 (FAD). Heterohelix reussi (Cushman. 1938) (FAWZ). Assigned age: latest Ceno-
manian to earliest Turonian.
Hedbergelltl (W.) aprka-Hedbergella (W.) baUica Z o n ~ : Marginotruncana undulata (Lehmann.
1963) (FAD). Hedbergella (Whiteinella) aprica (Loeblich & Tappan. 1961) (LAD). Hedbergella (Whitei-
nella) baltica Douglas & Rankin,l969 (LAD), Heterohelix moremani (Cushman. 1938) (LAWZ). Assigned
age: early to late Turonian.
Dicarinella primitiva Zone: Dicarinella primiliva (Dalbiez. 1955) (FAD). Guembelilria cenomana
(Keller. 1935) (LAD). Assigned age: latest Turonian.
Archaeoglobigerina cretacea-Dicarinellaprimitiva Zone: Archaeoglobigerina spp. (A. blowi Pes-
sagno. 1967, A. crelacea (d'Orbigny. 1840) ; (FAD). Dicarinella primiliva (Dalbiez. 1955) (LAD). Hedber-
gella (H.) ex gr. simplex (Morrow. 1934) (LAD). Assigned age: early to mid-Coniacian.
Dicarinelltl concavata-Marginotruncana sinuosa Zone: Marginotruncana sinuosa Porthault. 1970
(FAD). Dicarinella concavala (Brotzen. 1934) (FAD/LAD ; lR?). Hedbergella (H.) flandrini Porthault.
1970 (LAD), Marginotruncana renzi (Gandolfi. 1942) (LAWZ; late Coniacian). Marginotruncana undulata
(Lehmann. 1963) (LAWZ ; late Coniacian). Conlusolruncana ex gr.fornicata (plummer. 1931) (FAWZ;
Santonian). Ventilabrella auslinana Cushman. 1938 (FAWZ; Santonian). Assigned age: late Coniacian to
early-late Santonian.
Dicarinella asymetrica Zone: Dicarinella asymelrica (Sigal. 1952) (FAD/LAD; lR?), Hedbergel-
la (H.) ex gr. delrioensis (Carsey. 1926) (LAD). Assigned age: late Santonian.
Contusotruncana patelli/ormis-Globotruncanita elevata-stuarti/ormis plexus Zone :
Contusotruncanapatelliformis (Gandolfi, 1955) (FAD). Globotruncanitaelevata (Brotzen. 1934) - G. stuar-
tiformis (Dalbiez, 1955) plexus (FAD/LAD; lR?). Globotruncanella havanensis (Voorwijk. 1937) (FAD).
Globotruncanita stuartiformis 5.5. (FAD), Heterohelix reussi (Cushman) (LAWZ). Assigned age: early
Campanian.
Globotruncana orientaUs-Globotruncana ventricosa Zone: Globotruncana linneiana (d'Orbigny.
1839) (FAD). G. bu/loides Vogler, 1941 (FAD). G. orientalis El Naggar. 1966 (FAD). G. ventricosa White.
1928 (FAD). Globotruncanita subspinosa (pessagno. 1960) (FAD). Planoglobu/ina varians (Rzehak. 1895)
(FAD). Pseudotextularia elegans (Rzehak, 1891) (FAD). Rugoglobigerina ex gr. rugosa (plummer. 1926)
(FAD). Globotruncanita stuarti (de Lapparent. 1918) (FAD). Assigned age: early-late Campanian.
Contusotruncana ex gr. jornicata-Globotruncana linneiana Zone: Pseudotextularia browni
Masters. 1976 (FAD). Pseudoguembelina palpebra Bronnimann & Brown, 1953 (FAD), Contusotruncana
ex gr. fornicata (plummer. 1931) (LAD), Globotruncana bu/loides Vogler, 1941 (LAD). G. linneiana
(d' Orbigny. 1839) (LAD). Globotruncanita subspinosa (pessagno. 1960) (LAD). Gansserina gansseri
(Bolli. 1951) (FAWZ). Globotruncana aegyptiaca Nakkady, 1950 (FAWZ), Globotruncanella petaloidea
(Gandolfi. 1955) (FAWZ). Assigned age: early Maastrichtian.
Gansserinagansseri-Globotruncanitastuarti/ormisZone: Contusotruncanapatellijormis (Gandol-
fi, 1955) (LAD). Globotruncanita pettersi (Gandolfi. 1955) (FAD), Guembelitria cretacea Cushman. 1933
(FAD). Gansserina gansseri (Bolli. 1951) (LAD). Globotruncana rosetta (Carsey. 1926) (LAD). G.
ventricosaWhite. 1928(LAD). Globotruncanita stuartiformis (Dalbiez. 1955) S.s. (LAD). Assignedage: late
Maastrichtian.
Contusotruncana contusa-Globotruncana aegyptiaca Zone: Contusotruncana contusa (Cushman,
1926) (LFAD/LLAD ; lR). Globotruncana aegyptiaca Nakkady. 1950 (LAD). P/anoglobu/ina varians
254
Rotaliporabrol%eni Zone: Rotaliporabrotzeni (Sigal. 1948) (FAD/LAD; lR). Assignedage: earliest
Cenomanian.
Praeglobotruncana delrioensis-Rotalipora appenninica Zone :Hedbergella (Favusella) washiten-
sis (Carsey. 1926) (LAD). Praeglobotruncana delrioensis (plummer. 1931) (LAD). Rotalipora appennini-
ca (Reoz. 1936) (LAD). Assigned age: early to mid-Cenomanian.
Hedbergella (Whiteinelltl) baltica-Hedbergella (W.) brittonensis Zone: Hedbergella (Whiteinella)
baltica Douglas & Rankin. 1969 (FAD). Hedbergella (Whiteinella) brittonensis Loeblich & Tappan. 1961
(FAD). Assigned age: mid to late Cenomanian.
Hedbergelltl (W.)aprka-Globigerinelloides bentonensis Zone: Hedbergella (Whiteinella) aprica
(Loeblich & Tappan. 1961) (FAD), Globigerinelloides bentonensis (Morrow. 1934) (LAD). Assigned age:
late Cenomanian.
Hedbergella (W.) archaeocretacea-Heterohelix reussi Zone: Hedbergella (Whiteinella) archaeo-
eretacea Pessagno. 1967 (FAD). Heterohelix reussi (Cushman. 1938) (FAWZ). Assigned age: latest Ceno-
manian to earliest Turonian.
Hedbergelltl (W.) aprka-Hedbergella (W.) baUica Z o n ~ : Marginotruncana undulata (Lehmann.
1963) (FAD). Hedbergella (Whiteinella) aprica (Loeblich & Tappan. 1961) (LAD). Hedbergella (Whitei-
nella) baltica Douglas & Rankin,l969 (LAD), Heterohelix moremani (Cushman. 1938) (LAWZ). Assigned
age: early to late Turonian.
Dicarinella primitiva Zone: Dicarinella primiliva (Dalbiez. 1955) (FAD). Guembelilria cenomana
(Keller. 1935) (LAD). Assigned age: latest Turonian.
Archaeoglobigerina cretacea-Dicarinellaprimitiva Zone: Archaeoglobigerina spp. (A. blowi Pes-
sagno. 1967, A. erelacea (d'Orbigny. 1840) ; (FAD). Dicarinella primiliva (Dalbiez. 1955) (LAD). Hedber-
gella (H.) ex gr. simplex (Morrow. 1934) (LAD). Assigned age: early to mid-Coniacian.
Dicarinelltl concavata-Marginotruncana sinuosa Zone: Marginotruncana sinuosa Porthault. 1970
(FAD). Dicarinella concavala (Brotzen. 1934) (FAD/LAD ; lR?). Hedbergella (H.) flandrini Porthault.
1970 (LAD), Marginotruncana renzi (Gandolfi. 1942) (LAWZ; late Coniacian).Marginotruncana undulata
(Lehmann. 1963) (LAWZ ; late Coniacian). Conlusolruncana ex gr.fornicata (plummer. 1931) (FAWZ;
Santonian). Ventilabrella auslinana Cushman. 1938 (FAWZ; Santonian). Assigned age: late Coniacian to
early-late Santonian.
Dicarinella asymetrica Zone: Dicarinella asymelrica (Sigal. 1952) (FAD/LAD; lR?), Hedbergel-
la (H.) ex gr. delrioensis (Carsey. 1926) (LAD). Assigned age: late Santonian.
Contusotruncana patellijormis-Globotruncanita elevata-stuartijormis plexus Zone :
Contusotruncanapatelliformis (Gandolfi, 1955) (FAD). Globotruncanitaelevata (Brotzen. 1934) - G. stuar-
tiformis (Dalbiez, 1955) plexus (FAD/LAD; lR?). Globotruncanella havanensis (Voorwijk. 1937) (FAD).
Globotruncanita stuartiformis S.s. (FAD), Heterohelix reussi (Cushman) (LAWZ). Assigned age: early
Campanian.
Globotruncana orientaUs-Globotruncana ventricosa Zone: Globotruncana linneiana (d'Orbigny.
1839) (FAD). G. bu/loides Vogler, 1941 (FAD). G. orienta/is El Naggar. 1966 (FAD). G. ventricosa White.
1928 (FAD). Globotruncanita subspinosa (pessagno. 1960) (FAD). Pianoglobu/ina varians (Rzehak. 1895)
(FAD). Pseudotextularia elegans (Rzehak, 1891) (FAD). Rugoglobigerina ex gr. rugosa (plummer. 1926)
(FAD). Globotruncanita stuarti (de Lapparent. 1918) (FAD). Assigned age: early-late Campanian.
Contusotruncana ex gr. jornicata-Globotruncana linneiana Zone: Pseudotextularia browni
Masters. 1976 (FAD). Pseudoguembelina palpebra Bronnimann & Brown, 1953 (FAD), Contusotruncana
ex gc. fornicata (plummer. 1931) (LAD), Globotruncana bu/loides Vogler, 1941 (LAD). G. /inneiana
(d'Orbigny. 1839) (LAD). Globotruncanita subspinosa (pessagno. 1960) (LAD). Gansserina gansseri
(Bolli. 1951) (FAWZ). Globotruncana aegyptiaca Nakkady, 1950 (FAWZ), Globotruncanella petaloidea
(Gandolfi. 1955) (FAWZ). Assigned age: early Maastrichtian.
Gansserinagansseri-GlobotruncanitastuartiformisZone: Contusotruncanapatelliformis (Gandol-
fi, 1955) (LAD). Globotruncanita pettersi (Gandolfi. 1955) (FAD), Guembe/itria eretacea Cushman. 1933
(FAD). Gansserina gansseri (Bolli. 1951) (LAD). Globotruncana rosetta (Carsey. 1926) (LAD). G.
ventricosaWhite. 1928(LAD). Globotruncanita stuartiformis (Dalbiez. 1955) S.s. (LAD). Assignedage: late
Maastrichtian.
Contusotruncana contusa-Globotruncana aegyptiaca Zone: Contusotruncana contusa (Cushman,
1926) (LFAD/LLAD ; lR). Globotruncana aegyptiaca Nakkady. 1950 (LAD). P/anoglobu/ina varians
254
(Rzehak, 1895) (TR), and Globotruncana spp. (LAD), Rugoglobigerina spp. (LAD), Archaeoglobigerina
spp. (LAD), Globotruncanita spp. (LAD), Globigerinelloides spp. (LAD). Assigned age: latest Maastrieh-
tian.
BENTHIC FORAMINIFERIDA
Ungulogavelinella ciryi-Lingulonodosaria nodosaria-Marginulina ex gr. aequivoca Zone: Cate-
gory: Assemblage-zone. Lingulogavelinella ciryi Malapris-Bizouard, 1967 (OWZ), Lingulonodosaria no-
dosaria (Reuss, 1863) (OWZ), Marginulina ex gr. aequivocaReuss, 1863 (OWZ). Assigned age: late Aptian.
Ammobaculites cr. coprolithiformis-Buccicrenata hedbergi-Haplophragmium lueckei Zone:
Category: Assemblage-zone. Ammobaculites cf. coprolithiformis (Sehwager, 1867) (OWZ), Buccicrenata
hedbergi (Mayne, 1953) (OWZ), Haplophragmium lueckei (Cushman & Hedberg, 1941) (OWZ). Assigned
age: late Aptian.
Lenticulina ex gr. ex gr. nodosa Zone: Lenticulina ex gr. subangulata
(Reuss, 1863) (FAD),Budashevaella nonioninoides (Reuss, 1863) (FAD), Lenticulina ex gr. nodosa (Reuss,
1863) (LAD). Assigned age: late Aptian.
Epistominacarpenteri-Gaudryinopsisfl1iformisZone: Epistomina carpenteri (Reuss, 1863) (FAD),
E.chapmanitenDam, 1948 (FAD),Gaudryinopsiscf.gradata(Berthelin, 1880) (FAD),Pseudogaudryinella/
Spiroplectinata ex gr. dividens (Grabert, 1959) (FAD), Buccicrenata hedbergi (Maync, 1953) (LAD),
Gaudryinopsis filiformis (Berthelin, 1880) (LAD). Assigned age: earliest Albian.
Osangularia ex gr.dividensZone: Osangularia
schloenbachi (Reuss, 1863) (FAD), Pseudogaudryinella/Spiroplectinata ex gr. dividens (Grabert, 1959)
(LAD), Epistomina chapmani ten Dam, 1948 (LAD), Gavelinella barremiana Beuenstaedt, 1952 - G.
flandrini Moullade, 1960 plexus (LAD), Budashevaella nonioninoides (Reuss, 1863) (LAD), Marssonella
ozawai Cushman, 1936 (LAD), Tristix ex gr. excavata (Reuss, 1863) (FAWZ). Assigned age: early Albian.
Epistomina spinulifera-Epistomina carpenteri Zone: Epistomina spinulifera (Reuss, 1863) (FAD),
Cassidella ex gr. viscidus (Khan, 1950) (FAD), Epistomina carpenteri (Reuss, 1863) (LAD), Globorotali-
tes eL multisepta (Brotzen, 1936) (LAD). Assigned age: mid-Albian.
Gavelinella berthelini-plummerae-reussi plexus-Gaudryinopsis cf. gradata Zone : Osangularia
schloenbachi (Reuss, 1863) (LAD), Quasispiroplectammina ex gr. alexanderi (Lalieker, 1935) (LAD), Q. ex
gr. goodlandana (Lalicker, 1935) (LAD), Q. linki (petri, 1962) (LAD), Gaudryinopsis ef. gradata (Berthe-
lin, 1880) (LAWZ), Gavelinella berthelini (Keller, 1935) - G. plummerae (Tappan, 1940) - G. reussi (Khan,
1950) plexus (FAWZ), Textularia ex gr. subconica Franke, 1928 (FAWZ). Assigned age: mid to late Albian.
Neobulimina subcretacea-Saracenaria cf. crassicosta Zone: Neobulimina subcretacea Cushman,
1936 (FAD), Saracenaria ef. crassicosta Eiehenberg, 1933 (FAD). Assigned age: late Albian.
Neobulimina subcretacea-Tristix. ex gr. excavata Zone: Ramulina tetrahedralis Ludbrook, 1966
(FADILAD; TR?), Neobulimina subcretacea Cushman, 1936 (LAD), Tristix ex gr. excavata (Reuss, 1863)
(LAD), Lenticulina ex gr. subangulata (Reuss, 1863) (LAD). Assigned age: latest Albian.
Cibicides sp. A-Planularia complanata Zone: Cibicides sp. A (FAD), Planularia complanata
(Reuss, 1845) (FAD), Praebulimina ex gr. nannina (Tappan, 1940) (FAD), Saracenaria ef. crassicosta
Eichenberg, 1933 (LAWZ). Assigned age: earliest Cenomanian.
Lingulogavelinella(?) cr. thalmanniformis-Spiroloculina cretacea Zone: Lingulogavelinella(?) ef.
thalmanniformis (plotnikova, 1962) (FAD), Lingulogavelinella tormarpensis (Brotzen, 1942) (FAD),
Spiroloculina cretaceaReuss, 1854 (FADILAD ; TR?), Spiroloculina sp. A(FAD), Gavelinellasp. A(FAD).
Assigned age: early Cenomanian.
Nodosaria ex gr. obscura-Cibicides sp. A Zone: Nodosaria ex gr. obscura Reuss, 1845 (FAD),
Cibieides sp. A (LAD). Assigned age: mid to late Cenomanian.
Gabonita levis-Discammina sp. A Zone: Gabonita levis (de Klasz, Marie & Recat, 1961) (FAD),
Ammobaculites impexus Eieher, 1965 (FADILAD; TR?), Ammomarginulina paterella Eieher, 1967 (FAD/
LAD; TR?), Discammina sp. A (FADILAD ; TR?). Assigned age: late Cenomanian.
Gabonita obesa - Gabonita levis Zone: Gabonita obesa (de Klasz, Marie & Rerat, 1961) (FAD/
LAD; TR?), Buliminella sp. A (FAD), Cassidella ex gr. viscidus (Khan, 1950) (LAD), Gabonita levis (de
Klasz, Marie & Rerat, 1961) (AC). Assigned age: latest Cenomanian-earliest Turonian.
255
(Rzehak, 1895) (TR), and Globotruncana spp. (LAD), Rugoglobigerina spp. (LAD), Archaeoglobigerina
spp. (LAD), Globotruncanita spp. (LAD), Globigerinelloides spp. (LAD). Assigned age: latest Maastrich-
tian.
BENTHIC FORAMINIFERIDA
Ungulogavelinella ciryi-Lingulonodosaria nodosaria-Marginulina ex gr. aequivoca Zone: Cate-
gory: Assemblage-zone. Lingulogavelinella ciryi Malapris-Bizouard, 1967 (OWZ), Lingulonodosaria no-
dosaria (Reuss, 1863) (OWZ), Marginulina ex gr. aequivocaReuss, 1863 (OWZ). Assigned age: late Aptian.
Ammobaculites cr. coprolithijormis-BuccicrenatIJ hedbergi-Haplophragmium lueckei Zone:
Category: Assemblage-zone. Ammobaculites cr. coprolithijormis (Schwager, 1867) (OWZ), Buccicrenata
hedbergi (Mayne, 1953) (OWZ), Haplophragmium lueckei (Cushman & Hedberg, 1941) (OWZ). Assigned
age: late Aptian.
Lenticulina ex gr. ex gr. nodosa Zone: Lenticulina ex gr. subangulata
(Reuss, 1863) (FAD),Budashevaella nonioninoides (Reuss, 1863) (FAD), Lenticulina ex gr. nodosa (Reuss,
1863) (LAD). Assigned age: late Aptian.
Epistominacarpenteri-Gaudryinopsisfl1ijormisZone: Epistomina carpenteri (Reuss, 1863) (FAD),
E.chapmanitenDam, 1948 (FAD),Gaudryinopsiscf.gradata(Berthelin, 1880) (FAD),Pseudogaudryinella/
Spiroplectinata ex gr. dividens (Grabert, 1959) (FAD), Buccicrenata hedbergi (Mayne, 1953) (LAD),
Gaudryinopsis[ili/ormis (Berthelin, 1880) (LAD). Assigned age: earliest Albian.
Osangularia ex gr.dividensZone: Osangularia
schloenbachi (Reuss, 1863) (FAD), Pseudogaudryinella/Spiroplectinata ex gr. dividens (Grabert, 1959)
(LAD), Epistomina chapmani ten Dam, 1948 (LAD), Gavelinella barremiana Beuenstaedt, 1952 - G.
[landrini Moullade, 1960 plexus (LAD), Budashevaella nonioninoides (Reuss, 1863) (LAD), Marssonella
ozawai Cushman, 1936 (LAD), Tristix ex gr. excavata (Reuss, 1863) (FAWZ). Assigned age: early Albian.
Epistomina spinulijera-Epistomina carpenteri Zone: Epistomina spinuli/era (Reuss, 1863) (FAD),
Cassidella ex gr. viscidus (Khan, 1950) (FAD), Epistomina carpenteri (Reuss, 1863) (LAD), Globorotali-
tes cf. multisepta (Brotzen, 1936) (LAD). Assigned age: mid-Albian.
Gavelinella berthelini-plummerae-reussi plexus-Gaudryinopsis cr. gradatIJ Zone : Osangularia
schloenbachi (Reuss, 1863) (LAD), Quasispiroplectammina ex gr. alexanderi (Lalicker, 1935) (LAD), Q. ex
gr. goodlandana (Lalicker, 1935) (LAD), Q. linki (petri, 1962) (LAD), Gaudryinopsis cf. gradata (Berthe-
lin, 1880) (LAWZ), Gavelinella berthelini (Keller, 1935) - G. plummerae (Tappan, 1940) - G. reussi (Khan,
1950) plexus (FAWZ), Textularia ex gr. subconica Franke, 1928 (FAWZ). Assigned age: mid to late Albian.
Neobulimina subcretacea-Saracenaria cr. crassicostIJ Zone: Neobulimina subcretacea Cushman,
1936 (FAD), Saracenaria cf. crassicosta Eichenberg, 1933 (FAD). Assigned age: late Albian.
Neobulimina subcretacea-Tristix. ex gr. excavata Zone: Ramulina tetrahedralis Ludbrook, 1966
(FADILAD; TR?), Neobulimina subcretacea Cushman, 1936 (LAD), Trista ex gr. excavata (Reuss, 1863)
(LAD), Lenticulina ex gr. subangulata (Reuss, 1863) (LAD). Assigned age: latest Albian.
Cibicides sp. A-Planularia complanata Zone: Cibicides sp. A (FAD), Planularia complanata
(Reuss, 1845) (FAD), Praebulimina ex gr. nannina (Tappan, 1940) (FAD), Saracenaria cf. crassicosta
Eichenberg, 1933 (LAWZ). Assigned age: earliest Cenomanian.
Lingulogavelinella(?) cr. thalmannijormis-Spiroloculina cretacea Zone: Lingulogavelinella(?) cf.
thalmanni/ormis (plotnikova, 1962) (FAD), Lingulogavelinella tormarpensis (Bratzen, 1942) (FAD),
Spiroloculina cretaceaReuss, 1854 (FADILAD ; TR?), Spiroloculina sp. A(FAD), Gavelinellasp. A(FAD).
Assigned age: early Cenomanian.
Nodosaria ex gr. obscura-Cibicides sp. A Zone: Nodosaria ex gr. obscura Reuss, 1845 (FAD),
Cibicides sp. A (LAD). Assigned age: mid to late Cenomanian.
Gabonita levis-Discammina sp. A Zone: Gabonita levis (de Klasz, Marie & Recat, 1961) (FAD),
Ammobaculites impexus Eicher, 1965 (FADILAD; TR?), Ammomarginulina paterella Eicher, 1967 (FAD/
LAD; TR?), Discammina sp. A (FADILAD ; TR?). Assigned age: late Cenomanian.
Gabonita obesa - GabonitIJ levis Zone: Gabonita obesa (de Klasz, Marie & Rerat, 1961) (FAD/
LAD; TR?), Buliminella sp. A (FAD), Cassidella ex gr. viscidus (Khan, 1950) (LAD), Gabonita levis (de
Klasz, Marie & Rerat, 1961) (AC). Assigned age: latest Cenomanian-earliest Turonian.
255
(Rzehak, 1895) (TR), and Globotruncana spp. (LAD), Rugoglobigerina spp. (LAD), Archaeoglobigerina
spp. (LAD), Globotruncanita spp. (LAD), Globigerinelloides spp. (LAD). Assigned age: latest Maastrieh-
tian.
BENTHIC FORAMINIFERIDA
Ungulogavelinella ciryi-Lingulonodosaria nodosaria-Marginulina ex gr. aequivoca Zone: Cate-
gory: Assemblage-zone. Lingulogavelinella ciryi Malapris-Bizouard, 1967 (OWZ), Lingulonodosaria no-
dosaria (Reuss, 1863) (OWZ), Marginulina ex gr. aequivocaReuss, 1863 (OWZ). Assigned age: late Aptian.
Ammobaculites cr. coprolithijormis-BuccicrenatIJ hedbergi-Haplophragmium lueckei Zone:
Category: Assemblage-zone. Ammobaculites cf. coprolithijormis (Sehwager, 1867) (OWZ), Buccicrenata
hedbergi (Mayne, 1953) (OWZ), Haplophragmium lueckei (Cushman & Hedberg, 1941) (OWZ). Assigned
age: late Aptian.
Lenticulina ex gr. ex gr. nodosa Zone: Lenticulina ex gr. subangulata
(Reuss, 1863) (FAD),Budashevaella nonioninoides (Reuss, 1863) (FAD), Lenticulina ex gr. nodosa (Reuss,
1863) (LAD). Assigned age: late Aptian.
Epistominacarpenteri-Gaudryinopsisfl1ijormisZone: Epistomina carpenteri (Reuss, 1863) (FAD),
E.chapmanitenDam, 1948 (FAD),Gaudryinopsiscf.gradata(Berthelin, 1880) (FAD),Pseudogaudryinella/
Spiroplectinata ex gr. dividens (Grabert, 1959) (FAD), Buccicrenata hedbergi (Maync, 1953) (LAD),
Gaudryinopsis filiformis (Berthelin, 1880) (LAD). Assigned age: earliest Albian.
Osangularia ex gr.dividensZone: Osangularia
schloenbachi (Reuss, 1863) (FAD), Pseudogaudryinella/Spiroplectinata ex gr. dividens (Grabert, 1959)
(LAD), Epistomina chapmani ten Dam, 1948 (LAD), Gavelinella barremiana Beuenstaedt, 1952 - G.
[landrini Moullade, 1960 plexus (LAD), Budashevaella nonioninoides (Reuss, 1863) (LAD), Marssonella
ozawai Cushman, 1936 (LAD), Tristix ex gr. excavata (Reuss, 1863) (FAWZ). Assigned age: early Albian.
Epistomina spinulijera-Epistomina carpenteri Zone: Epistomina spinulifera (Reuss, 1863) (FAD),
Cassidella ex gr. viscidus (Khan, 1950) (FAD), Epistomina carpenteri (Reuss, 1863) (LAD), Globorotali-
tes eL multisepta (Brotzen, 1936) (LAD). Assigned age: mid-Albian.
Gavelinella berthelini-plummerae-reussi plexus-Gaudryinopsis cf. gradatIJ Zone : Osangularia
schloenbachi (Reuss, 1863) (LAD), Quasispiroplectammina ex gr. alexanderi (Lalieker, 1935) (LAD), Q. ex
gr. goodlandana (Lalicker, 1935) (LAD), Q. linki (petri, 1962) (LAD), Gaudryinopsis ef. gradata (Berthe-
lin, 1880) (LAWZ), Gavelinella berthelini (Keller, 1935) - G. plummerae (Tappan, 1940) - G. reussi (Khan,
1950) plexus (FAWZ), Textularia ex gr. subconica Franke, 1928 (FAWZ). Assigned age: mid to late Albian.
Neobulimina subcretacea-Saracenaria cf. crassicostIJ Zone: Neobulimina subcretacea Cushman,
1936 (FAD), Saracenaria ef. crassicosta Eiehenberg, 1933 (FAD). Assigned age: late Albian.
Neobulimina subcretacea-Tristix. ex gr. excavata Zone: Ramulina tetrahedralis Ludbrook, 1966
(FADILAD; TR?), Neobulimina subcretacea Cushman, 1936 (LAD), Tristix ex gr. excavata (Reuss, 1863)
(LAD), Lenticulina ex gr. subangulata (Reuss, 1863) (LAD). Assigned age: latest Albian.
Cibicides sp. A-Planularia complanata Zone: Cibicides sp. A (FAD), Planularia complanata
(Reuss, 1845) (FAD), Praebulimina ex gr. nannina (Tappan, 1940) (FAD), Saracenaria ef. crassicosta
Eichenberg, 1933 (LAWZ). Assigned age: earliest Cenomanian.
Lingulogavelinella(?) cr. thalmannijormis-Spiroloculina cretacea Zone: Lingulogavelinella(?) ef.
thalmanniformis (plotnikova, 1962) (FAD), Lingulogavelinella tormarpensis (Brotzen, 1942) (FAD),
Spiroloculina cretaceaReuss, 1854 (FADILAD ; TR?), Spiroloculina sp. A(FAD), Gavelinellasp. A(FAD).
Assigned age: early Cenomanian.
Nodosaria ex gr. obscura-Cibicides sp. A Zone: Nodosaria ex gr. obscura Reuss, 1845 (FAD),
Cibieides sp. A (LAD). Assigned age: mid to late Cenomanian.
Gabonita levis-Discammina sp. A Zone: Gabonita levis (de Klasz, Marie & Recat, 1961) (FAD),
Ammobaculites impexus Eieher, 1965 (FADILAD; TR?), Ammomarginulina paterella Eieher, 1967 (FAD/
LAD; TR?), Discammina sp. A (FADILAD ; TR?). Assigned age: late Cenomanian.
Gabonita obesa - GabonitIJ levis Zone: Gabonita obesa (de Klasz, Marie & Rerat, 1961) (FAD/
LAD; TR?), Buliminella sp. A (FAD), Cassidella ex gr. viscidus (Khan, 1950) (LAD), Gabonita levis (de
Klasz, Marie & Rerat, 1961) (AC). Assigned age: latest Cenomanian-earliest Turonian.
255
Gabonitalevis-NodosariIJ ex gr. obsCllraZone: Gabonitacf. parva (de Klasz, Marie&Meijer, 1960)
(FAD), Gabonitalevis(de Klasz, Marie&Rerat, 1961) (LAD), Nodosariaexgr. obscuraReuss, 1845(LAD).
Assigned age : early to late Turonian.
ValvlllineriIJ sp. A-GavelinelllJ berthelini-plllmmerae-rellssi plexus Zone: Valvulineria sp. A
(FAD), Bolivina cf. incrassata Reuss, 1851 (FAD), Gavelinella berthelini (Keller, 1935) - G. plummerae
(Tappan, 1940) - G. reussi (Khan, 1950) plexus (LAD). Assigned age: latest Turonian.
GavelinelllJ sp. A-ValvlllineriIJ sp. A Zone: Gavelinella sp. A (LAD), Valvulineria sp. A (LAD),
Lingulogavelinel/a tormarpensis (Brotzen, 1942) (LAD), Turrispirillina ex gr. subconica Tappan, 1943
(LAD). Assigned age: early to mid-Coniacian.
Lenticlllina revolllta-Globorotalites spinea Zone: Lenticulina revoluta (Israelsky, 1955) (FAD/
LAWZ ; late Coniacian ; TR?), Gavelinopsis sp. A (FAD/LAWZ ; late Coniacian ; TR?),Globorotalites
spinea (Cushman, 1926) (FAD/LAD ; TR?), Eponides aracajuensis Petri, 1962 (FAD), Gavelinella ex gr.
beccariiformis (White, 1928) (FAD), G. correcta (Carsey, 1926) (FAD), G. sandidgei (Brotzen, 1936)
(FAD), Gaudryina laevigata Franke, 1914 (FAD), Pseudogaudryinel/a ex gc. capitosa (Cushman, 1933)
(FAD), Gavelinella spissocostata (Cushman, 1938 (FAWZ ; Santonian), Reussel/a ex gr. szajnochae
(Grzybowski, 1896) (FAWZ; Santonian), Dorothia ex gr. bul/eta (Carsey, 1926) (FAWZ; Santonian),Re-
curvoides ex gr. globulosa (Grzybowski, 1896) (FAWZ; Santonian). Assigned age: late Coniacian to early-
late Santonian.
Nllttallinella texana-Orthokarstenia clarki Zone: Nuttallinel/a texana (Cushman, 1938) (FAD),
Orthokarstenia clarki (Cushman & Campbell, 1936) (FAD), O. clavata (Chenouard, de Klasz & Meijer,
1960) (FAD), Bolivina ex gr. afra (Reyment, 1959) (FAD), Verneuilina cretacea Karrer, 1870 (FAD),
Glaphyrammina sp. A (FAD/LAD ; TR?). Assigned age: late Santonian.
Siphogenerinoides bramlettei-Rellssella ex gr. s1lJjnochae Zone: Siphogenerinoides bramlettei
Cushman, 1929 (FAD), Ellipsoglandulina velascoensis Cushman, 1926 (FAD), Gavelinella nacatochensis
(Cushman, 1938) (FAD), Globorotalites subconica (Morrow, 1934) (FAD), Gyroidinoides loetterlei (Tap-
pan, 1940) (FAD), Osangularia navarroana (Cushman, 1938) (FAD), Planulina taylorensis (Carsey, 1926)
(FAD), Hagenowella ex gr. subsphaerica (Reuss, 1846) (FAD), Recurvoides cf. subturbinata (Grzybowski,
1898) (FAD), Trochamminoidesflagleri Cushman & Hedberg, 1941 (FAD), Reussella ex gr. szajnochae
(Grzybowski, 1896) (LAD), Pullenia jarvisi Cushman, 1936 (LAD). Assigned age: early Campanian.
Lacosteina gouskovi-Orthokarstenia clavata Zone: Lacosteina gouskovi Marie, 1945 (FAD/LAD;
TR?), Ammoglobigerina ex gr. globigeriniformis (parker & Jones, 1865) (FAD), Epistomina supracretacea
ten Dam, 1948 (FAD/LAD; TR?), GavelinellamonterelensisMarie, 1941 (FAD),Karreriellaexgr. conversa
(Grzybowski, 1901) (FAD), Neoflabelina rugosa (d'Orbigny, 1840) (FAD/LAD ; TR?), Praebulimina
spinata(Cushman &Campbell,1935) (FAD), Osangularianavarroana (Cushman, 1938) (LAWZ),Planulina
taylorensis (Carsey, 1926) (LAWZ), Orthokarstenia clavata (Chenouard, de Klasz &Meijer, 1960) (LAD),
Gavelinella correcta (Carsey, 1926) (LAD), G.lorneiana (d'Orbigny, 1840) (LAD), G. sandidgei (Brotzen,
1936) (LAD), Globorotalites subconica (Morrow, 1934) (LAD), Orithostel/a ex gc. halfeldi (petri, 1962)
(FAD), Valvulineria amaraliPetri, 1962 (LAD),B. ex gr. afra (AC; at the upper limit). Assigned age: early
to late Campanian.
Cibicides ex gr. beaumontiana-Gyroidinoides nonionoides Zone: Ammobaculites cf.fragmentaria
Cushman, 1927(FAD), Cibicidesex gr. beaumontiana(d'Orbigny, 1840) (LAD), Gyroidinoidesnonionoides
(Bandy, 1951) (LAD), Neoflabelina reticulata (Reuss, 1851) (LAD), N. ex gr. pilulifera (Cushman &
Campbell, 1935) (LAWZ), Praebulimina ex gr.fang de Klasz, Magne &Rerat, 1963 (LAD), P. ex gc. prolixa
(Cushman & Parker, 1925) (LAD), P. spinata (Cushman & Campbell, 1935) (LAD). Assigned age: early
Maastrichtian.
Gyroidinoides loetterki-OsanglllariIJ velascoensis Zone: Gyroidinoides loetterlei (Tappan, 1940)
(LAD), Osangularia velascoensis (Cushman, 1925) (LAD), Nodogenerina stephensoni (Cushman, 1936)
(LAD), Praebulimina ex gr. bantu de Klasz, Magne & Rerat, 1963 (LAD), Ammobaculites cf.fragmentaria
Cushman, 1927 (LAD), RzehakinaflSsistomata (Grzybowski, 1901) (LAD), Verneuilina cretosa Cushman,
1933 (LAD). Assigned age: late Maastrichtian.
Orthokarstenia clarki-Praeblllimina kickapooensis Zone: Clavlllinoides ex gr. trilatera (Cushman,
1926) (FAD), Gaudryinopsis glabrata (Cushman, 1933) (FAD),Pseudoclavulina arenata (Cushman, 1933)
(FAD), Orthokarstenia clarki (Cushman &Campbell, 1936) (LAD), Buliminel/aex gc. colonensis Cushman
256
GabonitalevisNodosariIJ ex gr. obsCllraZone: Gabonitacf. parva (de Klasz, Marie& Meijer, 1960)
(FAD), Gabonitalevis(de Klasz, Marie& Rerat, 1961) (LAD), Nodosariaexgr. obseuraReuss, 1845(LAD).
Assigned age : early to late Turonian.
ValvlllineriIJ sp. AGavelinelllJ bertheliniplllmmeraerellssi plexus Zone: Valvulineria sp. A
(FAD), Bolivina cf. incrassata Reuss, 1851 (FAD), Gavelinella berthelini (Keller, 1935) - G. plummerae
(Tappan, 1940) - G. reussi (Khan, 1950) plexus (LAD). Assigned age: latest Turonian.
GavelinelllJ sp. AValvlllineriIJ sp. A Zone: Gavelinella sp. A (LAD), Valvulineria sp. A (LAD),
Lingulogavelinella tormarpensis (Brotzen, 1942) (LAD), Turrispirillina ex gr. subeoniea Tappan, 1943
(LAD). Assigned age: early to mid-Coniacian.
Lenticlllina revollltaGloborotalites spinea Zone: Lentieulina revoluta (Israelsky, 1955) (FAD/
LAWZ ; late Coniacian; TR?), Gavelinopsis sp. A (FAD/LAWZ ; late Coniacian; TR?),Globorotalites
spinea (Cushman, 1926) (FAD/LAD ; TR?), Eponides araeajuensis Petri, 1962 (FAD), Gavelinella ex gr.
beeeariijormis (White, 1928) (FAD), G. eorreeta (Carsey, 1926) (FAD), G. sandidgei (Brotzen, 1936)
(FAD), Gaudryina laevigata Franke, 1914 (FAD), Pseudogaudryinella ex gr. eapitosa (Cushman, 1933)
(FAD), Gavelinella spissoeostata (Cushman, 1938 (FAWZ ; Santonian), Reussella ex gr. szajnoehae
(Grzybowski, 1896) (FAWZ; Santonian), Dorothia ex gr. bulleta (Carsey, 1926) (FAWZ; Santonian),Re-
eurvoides ex gr. globulosa (Grzybowski, 1896) (FAWZ; Santonian). Assigned age: late Coniacian to early-
late Santonian.
Nllttallinella texanaOrthokarstenia clarki Zone: Nuttallinella texana (Cushman, 1938) (FAD),
Orthokarstenia clarki (Cushman & Campbell, 1936) (FAD), O. clavata (Chenouard, de Klasz & Meijer,
1960) (FAD), Bolivina ex gr. ajra (Reyment, 1959) (FAD), Verneuilina cretaeea Karrer, 1870 (FAD),
Glaphyrammina sp. A (FAD/LAD ; TR?). Assigned age: late Santonian.
Siphogenerinoides bramletteiRellssella ex gr. s1lJjnochae Zone: Siphogenerinoides bramlettei
Cushman, 1929 (FAD), Ellipsoglandulina velaseoensis Cushman, 1926 (FAD), Gavelinella naeatoehensis
(Cushman, 1938) (FAD), Globorotalites subeonica (Morrow, 1934) (FAD), Gyroidinoides loetterlei (Tap-
pan, 1940) (FAD), Osangularia navarroana (Cushman, 1938) (FAD), Planulina taylorensis (Carsey, 1926)
(FAD), Hagenowella ex gr. subsphaeriea (Reuss, 1846) (FAD), Reeurvoides cf. subturbinata (Grzybowski,
1898) (FAD), Trochamminoidesjlagleri Cushman & Hedberg, 1941 (FAD), Reussella ex gr. szajnoehae
(Grzybowski, 1896) (LAD), Pullenia jarvisi Cushman, 1936 (LAD). Assigned age: early Campanian.
Lacosteina gouskovi-Orthokarstenia clavata Zone: Laeosteina gouskovi Marie, 1945 (FAD/LAD;
TR?), Ammoglobigerina ex gr. globigerinijormis (parker & Jones, 1865) (FAD), Epistomina supracretaeea
ten Dam, 1948 (FAD/LAD; TR?), GavelinellamonterelensisMarie, 1941 (FAD),Karreriellaexgr. eonversa
(Grzybowski, 1901) (FAD), Neojlabelina rugosa (d'Orbigny, 1840) (FAD/LAD ; TR?), Praebulimina
spinata(Cushman & Campbell,1935) (FAD), Osangularianavarroana (Cushman, 1938) (LAWZ),Planulina
taylorensis (Carsey, 1926) (LAWZ), Orthokarstenia clavata (Chenouard, de Klasz & Meijer, 1960) (LAD),
Gavelinella eorreeta (Carsey, 1926) (LAD), G.lorneiana (d'Orbigny, 1840) (LAD), G. sandidgei (Brotzen,
1936) (LAD), Globorotalites subeoniea (Morrow, 1934) (LAD), Orithostella ex gr. halfeldi (petri, 1962)
(FAD), Valvulineria amaraliPetri, 1962 (LAD),B. ex gr. afra (AC; at the upper limit). Assigned age: early
to late Campanian.
Cibicides ex gr. beaumontiana-Gyroidinoides nonionoides Zone: Ammobaeulites cf.jragmentaria
Cushman, 1927(FAD), Cibicidesex gr. beaumontiana(d'Orbigny, 1840) (LAD), Gyroidinoidesnonionoides
(Bandy, 1951) (LAD), Neojlabelina retieulata (Reuss, 1851) (LAD), N. ex gr. pi/ulijera (Cushman &
Campbell, 1935) (LAWZ), Praebulimina ex gr.jang de Klasz, Magne & Rerat, 1963 (LAD), P. ex gr. prolixa
(Cushman & Parker, 1925) (LAD), P. spinata (Cushman & Campbell, 1935) (LAD). Assigned age: early
Maastrichtian.
Gyroidinoides loetterkiOsanglllariIJ velascoensis Zone: Gyroidinoides loetterlei (Tappan, 1940)
(LAD), Osangularia velaseoensis (Cushman, 1925) (LAD), Nodogenerina stephensoni (Cushman, 1936)
(LAD), Praebulimina ex gr. bantu de Klasz, Magne & Rerat, 1963 (LAD), Ammobaeulites cf.jragmentaria
Cushman, 1927 (LAD), Rzehakina[lSsistomata (Grzybowski, 1901) (LAD), Verneuilina eretosa Cushman,
1933 (LAD). Assigned age: late Maastrichtian.
Orthokarstenia clarkiPraeblllimina kickapooensis Zone: Clavlliinoides ex gr. tri/atera (Cushman,
1926) (FAD), Gaudryinopsis glabrata (Cushman, 1933) (FAD),Pseudoclavulina arenata (Cushman, 1933)
(FAD), Orthokarstenia clarki (Cushman & Campbell, 1936) (LAD), Buliminel/aex gr. eolonensis Cushman
256
GabonitalevisNodosariIJ ex gr. obsCllraZone: Gabonitacf. parva (de Klasz, Marie&Meijer, 1960)
(FAD), Gabonitalevis(de Klasz, Marie&Rerat, 1961) (LAD), Nodosariaexgr. obscuraReuss, 1845(LAD).
Assigned age : early to late Turonian.
ValvlllineriIJ sp. AGavelinelllJ bertheliniplllmmeraerellssi plexus Zone: Valvulineria sp. A
(FAD), Bolivina cf. incrassata Reuss, 1851 (FAD), Gavelinella berthelini (Keller, 1935) - G. plummerae
(Tappan, 1940) - G. reussi (Khan, 1950) plexus (LAD). Assigned age: latest Turonian.
GavelinelllJ sp. AValvlllineriIJ sp. A Zone: Gavelinella sp. A (LAD), Valvulineria sp. A (LAD),
Lingulogavelinella tormarpensis (Brotzen, 1942) (LAD), Turrispirillina ex gr. subconica Tappan, 1943
(LAD). Assigned age: early to mid-Coniacian.
Lenticlllina revollltaGloborotalites spinea Zone: Lenticulina revoluta (Israelsky, 1955) (FAD/
LAWZ ; late Coniacian ; TR?), Gavelinopsis sp. A (FAD/LAWZ ; late Coniacian ; TR?),Globorotalites
spinea (Cushman, 1926) (FAD/LAD ; TR?), Eponides aracajuensis Petri, 1962 (FAD), Gavelinella ex gr.
beccariijormis (White, 1928) (FAD), G. correcta (Carsey, 1926) (FAD), G. sandidgei (Brotzen, 1936)
(FAD), Gaudryina laevigata Franke, 1914 (FAD), Pseudogaudryinella ex gc. capitosa (Cushman, 1933)
(FAD), Gavelinella spissocostata (Cushman, 1938 (FAWZ ; Santonian), Reussella ex gr. szajnochae
(Grzybowski, 1896) (FAWZ; Santonian), Dorothia ex gr. bulleta (Carsey, 1926) (FAWZ; Santonian),Re-
curvoides ex gr. globulosa (Grzybowski, 1896) (FAWZ; Santonian). Assigned age: late Coniacian to early-
late Santonian.
Nllttallinella texanaOrthokarstenia clarki Zone: Nuttallinella texana (Cushman, 1938) (FAD),
Orthokarstenia clarki (Cushman & Campbell, 1936) (FAD), O. clavata (Chenouard, de Klasz & Meijer,
1960) (FAD), Bolivina ex gr. afra (Reyment, 1959) (FAD), Verneuilina cretacea Karrer, 1870 (FAD),
Glaphyrammina sp. A (FAD/LAD ; TR?). Assigned age: late Santonian.
Siphogenerinoides bramletteiRellssella ex gr. s1lJjnochae Zone: Siphogenerinoides bramlettei
Cushman, 1929 (FAD), Ellipsoglandulina velascoensis Cushman, 1926 (FAD), Gavelinella nacatochensis
(Cushman, 1938) (FAD), Globorotalites subconica (Morrow, 1934) (FAD), Gyroidinoides loetterlei (Tap-
pan, 1940) (FAD), Osangularia navarroana (Cushman, 1938) (FAD), Planulina taylorensis (Carsey, 1926)
(FAD), Hagenowella ex gr. subsphaerica (Reuss, 1846) (FAD), Recurvoides cf. subturbinata (Grzybowski,
1898) (FAD), Trochamminoidesjlagleri Cushman & Hedberg, 1941 (FAD), Reussella ex gr. szajnochae
(Grzybowski, 1896) (LAD), Pullenia jarvisi Cushman, 1936 (LAD). Assigned age: early Campanian.
Lacosteina gouskovi-Orthokarstenia clavata Zone: Lacosteina gouskovi Marie, 1945 (FAD/LAD;
TR?), Ammoglobigerina ex gr. globigerinijormis (parker & Jones, 1865) (FAD), Epistomina supracretacea
ten Dam, 1948 (FAD/LAD; TR?), GavelinellamonterelensisMarie, 1941 (FAD),Karreriellaexgr. conversa
(Grzybowski, 1901) (FAD), Neojlabelina rugosa (d'Orbigny, 1840) (FAD/LAD ; TR?), Praebulimina
spinata(Cushman &Campbell,1935) (FAD), Osangularianavarroana (Cushman, 1938) (LAWZ),Planulina
taylorensis (Carsey, 1926) (LAWZ), Orthokarstenia clavata (Chenouard, de Klasz &Meijer, 1960) (LAD),
Gavelinella correcta (Carsey, 1926) (LAD), G.lorneiana (d'Orbigny, 1840) (LAD), G. sandidgei (Brotzen,
1936) (LAD), Globorotalites subconica (Morrow, 1934) (LAD), Orithostella ex gc. halfeldi (petri, 1962)
(FAD), Valvulineria amaraliPetri, 1962 (LAD),B. ex gr. afra (AC; at the upper limit). Assigned age: early
to late Campanian.
Cibicides ex gr. beaumontiana-Gyroidinoides nonionoides Zone: Ammobaculites cf.fragmentaria
Cushman, 1927(FAD), Cibicidesex gr. beaumontiana(d'Orbigny, 1840) (LAD), Gyroidinoidesnonionoides
(Bandy, 1951) (LAD), Neojlabelina reticulata (Reuss, 1851) (LAD), N. ex gr. pilulijera (Cushman &
Campbell, 1935) (LAWZ), Praebulimina ex gr.fang de Klasz, Magne &Rerat, 1963 (LAD), P. ex gc. prolixa
(Cushman & Parker, 1925) (LAD), P. spinata (Cushman & Campbell, 1935) (LAD). Assigned age: early
Maastrichtian.
Gyroidinoides loetterkiOsanglllariIJ velascoensis Zone: Gyroidinoides loetterlei (Tappan, 1940)
(LAD), Osangularia velascoensis (Cushman, 1925) (LAD), Nodogenerina stephensoni (Cushman, 1936)
(LAD), Praebulimina ex gr. bantu de Klasz, Magne & Rerat, 1963 (LAD), Ammobaculites cf.fragmentaria
Cushman, 1927 (LAD), Rzehakina[lSsistomata (Grzybowski, 1901) (LAD), Verneuilina cretosa Cushman,
1933 (LAD). Assigned age: late Maastrichtian.
Orthokarstenia clarkiPraeblllimina kickapooensis Zone: Clavlllinoides ex gr. trilatera (Cushman,
1926) (FAD), Gaudryinopsis glabrata (Cushman, 1933) (FAD),Pseudoclavulina arenata (Cushman, 1933)
(FAD), Orthokarstenia clarki (Cushman &Campbell, 1936) (LAD), Buliminel/aex gc. colonensis Cushman
256
& Hedberg, 1930 (LAD), Neobulimina aspera (Cushman & Parker, 1940) (LAD), N. canadensis Cushman
& Wickenden, 1928 (LAD), Praebulimina kickapooensis (Cole, 1938) (LAD), P. reussi (Morrow, 1934)
(LAD), Pseudouvigerina plummerae Cushman, 1927 (LAD). Assigned age: latest Maastrichtian.
AMMONITES
EpichelonicerasDiadochoceras.EodouYilleiceras Zone: Category: Assemblage wne.Epichelonice-
ras sp. (OWZ), Diadochoceras sp. (OWZ), Eodouvilleiceras horridum(Riedel, 1937) (OWZ), Eodouvillei-
ceras sp. (OWZ), <<Dufrenoyia cf.justinae Hill, 1893 (OWZ). Assigned age: late Aptian.
Douvilleiceras Zone: Douvilleicerasex gr. mammillatum(Schlotheim, 1813) (FAD),Douvilleiceras
inaequinodum (Quenstedt, 1849) (OWZPR). Assigned age: early Albian.
Oxytropidoceras Zone: Oxytropidoceras buarquianum (White, 1887) (FAD), Oxytropidoceras spp.
(OWZ). Assigned age: mid Albian.
Elobiceras Zone: Elobiceras intermedium Spath, 1922 (FAD), Elobiceras sp. (OWZ). Assigned
age: early late Albian.
Mortoniceras Zone: Mortonicerascf.lastroensis Maury, 1937 (FAD), Mortoniceras spp. (OWZPR),
Neokentroceras cf. tectorium (White, 1887) (OWZPR). Assigned age: latest Albian.
Graysonites lozoiHypoturrilites betaitraensis Zone: Graysonites lozoi Young, 1958 (FAD),
Sharpeicerasvohipalense Collignon, 1964 (OWZPR), Hypoturrilites betaitraensis Klinger & Kennedy,
1978 (OWZPR), Stoliczkaia (Shumarinaia) africana Pervinquiere, 1907 (OWZPR), Forbesiceras brundret-
ti(Young, 1958) (OWZPR). Assigned age: early Cenomanian.
Acompsoceras spathiDunveganoceras Zone: Acompsoceras spathi Basse, 1940 (FAD), Acompso-
ceras aff. renevieri (Sharpe, 1857) (OWZPR), Dunveganoceras sp. (OWZPR), Euomphaloceras (E.) meri-
dionale (Stoliczka, 1864) (OWZ), Turrilites scheuchzerianus Bosc, 1801 (OWZ). Assigned age: early mid
Cenomanian.
AcanthocerasjukesbrowneiEucalycoceraspentagonumZone: Acanthocerasjukesbrownei (Spath,
1926) (FAD), Eucalycoceras pentagonum(Jukes-Browne & Hill, 1896) (OWZ). Assigned age : late midCe-
nomanian.
Pseudocalycoceras harpaxThomelites aft sornayi Zone: Pseudocalycoceras harpax (Stoliczka,
1864) (FAD), Thomelites aff. sornayi (Thomel, 1967) (OWZPR). Assigned age: early late Cenomanian.
Euomphaloceras septemseriatum Zone: Euomphaloceras septemseriatum (Cragin, 1893) (FAD),
Pseudaspidoceras pseudonodosoides (Choffat, 1899) (OWZ), Vascoceras gamai Choffat, 1899 (OWZ),
Thomasites gongilensis (Woods, 1911) (OWZ). Assigned age: latest Cenomanian.
Vascoceras harttiiPseudaspidocerasfooteanum Zone: Pseudotissotia nigeriensis (Woods, 1911)
(FAD), Pseudotissotia gabonensis Lombard, 1931 (OWZPR), Vascoceras hamii (Hyatt, 1870) (OWZPR),
Pseudaspidoceras footeanum (Stoliczka, 1864) (OWZ), Wrightoceras sp. (OWZ). Assigned age: latest
Cenomanian-earliest Turonian.
Watinoceras amudarienseKamerunoceras seitzi Zone: Watinoceras amudariense (Arkhangel' skij,
1916) (FAD), Kamerunoceras seitzi (Riedel, 1932) (OWZ), Mitonia reesidei (Maury, 1937) (OWZPR).
Assigned age: early Turonian.
. Mammites nodosoidesKamerunoceras turoniense Zone: Mammites nodosoides (Schltiter, 1871)
(FAD), Kamerunoceras turoniense (d'Orbigny, 1850) (OWZPR), Fagesia bomba Eck, 1909 (OWZPR),
Romaniceras deverianum (d'Orbigny, 1841) (OWZ). Assigned age: early-mid Turonian.
SubprionocyclusReesidites Zone : Subprionocyclus sp. (FAD), Reesidites sp. (OWZPR),
Paralenticeras leonhardianum (Karsten, 1858) (OWZ). Assigned age: late Turonian.
Bllrroisiceras (B.) onilahyense.Forresteria Zone: Barroisiceras (B.) onilahyense Basse, 1948
(FAD), Forresteria sp: (OWZ). Assigned age: early Coniacian.
Solgerites armatusPrionocycloceras lenti Zone : Solgerites armatus (Solger, 1904) (FAD),
Prionocycloceras lenti (Gerhardt, 1897) (OWZ), Prionocycloceras guayabanum(Steinmann, 1897) (OWZ),
Heterotissotia sp. (OWZ), Bostrychoceras indicum (Stoliczka, 1865) (OWZ), Peroniceras spp. (OWZ).
Assigned age: mid Coniacian.
257
& Hedberg, 1930 (LAD), Neobulimina aspera (Cushman & Parker, 1940) (LAD), N. canadensis Cushman
& Wickenden, 1928 (LAD), Praebulimina kickapooensis (Cole, 1938) (LAD), P. reussi (Morrow, 1934)
(LAD), Pseudouvigerina plummerae Cushman, 1927 (LAD). Assigned age: latest Maastrichtian.
AMMONITES
EpichelonicerasDiadochoceras.EodouYilleiceras Zone: Category: Assemblage wne.Epichelonice-
ras sp. (OWZ), Diadochoceras sp. (OWZ), Eodouvilleiceras horridum(Riedel, 1937) (OWZ), Eodouvillei-
ceras sp. (OWZ), (<Dufrenoyia cf.justinae Hill, 1893 (OWZ). Assigned age: late Aptian.
Douvilleiceras Zone: Douvilleicerasex gr. mammillatum(Schlotheim, 1813) (FAD),Douvilleiceras
inaequinodum (Quenstedt, 1849) (OWZPR). Assigned age: early Albian.
Oxytropidoceras Zone: Oxytropidoceras buarquianum (White, 1887) (FAD), Oxytropidoceras spp.
(OWZ). Assigned age: mid Albian.
Elobiceras Zone: Elobiceras intermedium Spath, 1922 (FAD), Elobiceras sp. (OWZ). Assigned
age: early late Albian.
Mortoniceras Zone: Mortonicerascf.lastroensis Maury, 1937 (FAD), Mortoniceras spp. (OWZPR),
Neokentroceras cf. tectorium (White, 1887) (OWZPR). Assigned age: latest Albian.
Graysonites lozoiHypoturrilites betaitraensis Zone: Graysonites lozoi Young, 1958 (FAD),
Sharpeicerasvohipalense Collignon, 1964 (OWZPR), Hypoturrilites betaitraensis Klinger & Kennedy,
1978 (OWZPR), Stoliczkaia (Shumarinaia) africana Pervinquiere, 1907 (OWZPR), Forbesiceras brundret-
ti(Young, 1958) (OWZPR). Assigned age: early Cenomanian.
Acompsoceras spathiDunveganoceras Zone: Acompsoceras spathi Basse, 1940 (FAD),Acompso-
ceras aff. renevieri (Sharpe, 1857) (OWZPR), Dunveganoceras sp. (OWZPR), Euomphaloceras (E.) meri-
dionale (Stoliczka, 1864) (OWZ), Turrilites scheuchzerianus Bose, 1801 (OWZ). Assigned age: early mid
Cenomanian.
AcanthocerasjukesbrowneiEucalycoceraspentagonumZone: Acanthocerasjukesbrownei (Spath,
1926) (FAD), Eucalycoceras pentagonum(Jukes-Browne & Hill, 1896) (OWZ). Assigned age : late midCe-
nomanian.
Pseudocalycoceras harpaxThomelites aft sornayi Zone: Pseudocalycoceras harpax (Stoliczka,
1864) (FAD), Thomelites aff. sornayi (Thomel, 1967) (OWZPR). Assigned age: early late Cenomanian.
Euomphaloceras septemseriatum Zone: Euomphaloceras septemseriatum (Cragin, 1893) (FAD),
Pseudaspidoceras pseudonodosoides (Choffat, 1899) (OWZ), Vascoceras gamai Choffat, 1899 (OWZ),
Thomasites gongilensis (Woods, 1911) (OWZ). Assigned age: latest Cenomanian.
Vascoceras harttiiPseudaspidoceras!ooteanum Zone: Pseudotissotia nigeriensis (Woods, 1911)
(FAD), Pseudotissotia gabonensis Lombard, 1931 (OWZPR), Vascoceras hamii (Hyatt, 1870) (OWZPR),
Pseudaspidoceras footeanum (Stoliczka, 1864) (OWZ), Wrightoceras sp. (OWZ). Assigned age: latest
Cenomanian-earliest Turonian.
Watinoceras amudarienseKamerunoceras seitzi Zone: Watinoceras amudariense (Arkhangel' skij,
1916) (FAD), Kamerunoceras seitzi (Riedel, 1932) (OWZ), Mitonia reesidei (Maury, 1937) (OWZPR).
Assigned age: early Turonian.
. Mammites nodosoidesKamerunoceras turoniense Zone: Mammites nodosoides (SchlUter, 1871)
(FAD), Kamerunoceras turoniense (d'Orbigny, 1850) (OWZPR), Fagesia bomba Eck, 1909 (OWZPR),
Romaniceras deverianum (d'Orbigny, 1841) (OWZ). Assigned age: early-mid Turonian.
SubprionocyclusReesidites Zone : Subprionocyclus sp. (FAD), Reesidites sp. (OWZPR),
Paralenticeras leonhardianum (Karsten, 1858) (OWZ). Assigned age: late Turonian.
Bllrroisiceras (B.) onilahyense.Forresteria Zone: Barroisiceras (B.) onilahyense Basse, 1948
(FAD), Forresteria sp: (OWZ). Assigned age: early Coniacian.
Solgerites armatusPrwnocycloceras lenti Zone : Solgerites armatus (Solger, 1904) (FAD),
Prionocycloceras lenti (Gerhardt, 1897) (OWZ), Prionocycloceras guayabanum(Steinmann, 1897) (OWZ),
Heterotissotia sp. (OWZ), Bostrychoceras indicum (Stoliczka, 1865) (OWZ), Peroniceras spp. (OWZ).
Assigned age: mid Coniacian.
257
& Hedberg, 1930 (LAD), Neobulimina aspera (Cushman & Parker, 1940) (LAD), N. canadensis Cushman
& Wickenden, 1928 (LAD), Praebulimina kickapooensis (Cole, 1938) (LAD), P. reussi (Morrow, 1934)
(LAD), Pseudouvigerina plummerae Cushman, 1927 (LAD). Assigned age: latest Maastrichtian.
AMMONITES
EpichelonicerasDiadochoceras.EodouYilleiceras Zone: Category: Assemblage wne.Epichelonice-
ras sp. (OWZ), Diadochoceras sp. (OWZ), Eodouvilleiceras horridum(Riedel, 1937) (OWZ), Eodouvillei-
ceras sp. (OWZ), <<Dufrenoyia cf.justinae Hill, 1893 (OWZ). Assigned age: late Aptian.
Douvilleiceras Zone: Douvilleicerasex gr. mammillatum(Schlotheim, 1813) (FAD),Douvilleiceras
inaequinodum (Quenstedt, 1849) (OWZPR). Assigned age: early Albian.
Oxytropidoceras Zone: Oxytropidoceras buarquianum (White, 1887) (FAD), Oxytropidoceras spp.
(OWZ). Assigned age: mid Albian.
Elobiceras Zone: Elobiceras intermedium Spath, 1922 (FAD), Elobiceras sp. (OWZ). Assigned
age: early late Albian.
Mortoniceras Zone: Mortonicerascf.lastroensis Maury, 1937 (FAD), Mortoniceras spp. (OWZPR),
Neokentroceras cf. tectorium (White, 1887) (OWZPR). Assigned age: latest Albian.
Graysonites lozoiHypoturrilites betaitraensis Zone: Graysonites lozoi Young, 1958 (FAD),
Sharpeicerasvohipalense Collignon, 1964 (OWZPR), Hypoturrilites betaitraensis Klinger & Kennedy,
1978 (OWZPR), Stoliczkaia (Shumarinaia) africana Pervinquiere, 1907 (OWZPR), Forbesiceras brundret-
ti(Young, 1958) (OWZPR). Assigned age: early Cenomanian.
Acompsoceras spathiDunveganoceras Zone: Acompsoceras spathi Basse, 1940 (FAD),Acompso-
ceras aff. renevieri (Sharpe, 1857) (OWZPR), Dunveganoceras sp. (OWZPR), Euomphaloceras (E.) meri-
dionale (Stoliczka, 1864) (OWZ), Turrilites scheuchzerianus Bosc, 1801 (OWZ). Assigned age: early mid
Cenomanian.
AcanthocerasjukesbrowneiEucalycoceraspentagonumZone: Acanthocerasjukesbrownei (Spath,
1926) (FAD), Eucalycoceras pentagonum(Jukes-Browne & Hill, 1896) (OWZ). Assigned age : late midCe-
nomanian.
Pseudocalycoceras harpaxThomelites aft sornayi Zone: Pseudocalycoceras harpax (Stoliczka,
1864) (FAD), Thomelites aff. sornayi (Thomel, 1967) (OWZPR). Assigned age: early late Cenomanian.
Euomphaloceras septemseriatum Zone: Euomphaloceras septemseriatum (Cragin, 1893) (FAD),
Pseudaspidoceras pseudonodosoides (Choffat, 1899) (OWZ), Vascoceras gamai Choffat, 1899 (OWZ),
Thomasites gongilensis (Woods, 1911) (OWZ). Assigned age: latest Cenomanian.
Vascoceras harttiiPseudaspidocerasfooteanum Zone: Pseudotissotia nigeriensis (Woods, 1911)
(FAD), Pseudotissotia gabonensis Lombard, 1931 (OWZPR), Vascoceras hamii (Hyatt, 1870) (OWZPR),
Pseudaspidoceras footeanum (Stoliczka, 1864) (OWZ), Wrightoceras sp. (OWZ). Assigned age: latest
Cenomanian-earliest Turonian.
Watinoceras amudarienseKamerunoceras seitzi Zone: Watinoceras amudariense (Arkhangel' skij,
1916) (FAD), Kamerunoceras seitzi (Riedel, 1932) (OWZ), Mitonia reesidei (Maury, 1937) (OWZPR).
Assigned age: early Turonian.
. Mammites nodosoidesKamerunoceras turoniense Zone: Mammites nodosoides (Schltiter, 1871)
(FAD), Kamerunoceras turoniense (d'Orbigny, 1850) (OWZPR), Fagesia bomba Eck, 1909 (OWZPR),
Romaniceras deverianum (d'Orbigny, 1841) (OWZ). Assigned age: early-mid Turonian.
SubprionocyclusReesidites Zone : Subprionocyclus sp. (FAD), Reesidites sp. (OWZPR),
Paralenticeras leonhardianum (Karsten, 1858) (OWZ). Assigned age: late Turonian.
Bllrroisiceras (B.) onilahyense.Forresteria Zone: Barroisiceras (B.) onilahyense Basse, 1948
(FAD), Forresteria sp: (OWZ). Assigned age: early Coniacian.
Solgerites armatusPrionocycloceras lenti Zone : Solgerites armatus (Solger, 1904) (FAD),
Prionocycloceras lenti (Gerhardt, 1897) (OWZ), Prionocycloceras guayabanum(Steinmann, 1897) (OWZ),
Heterotissotia sp. (OWZ), Bostrychoceras indicum (Stoliczka, 1865) (OWZ), Peroniceras spp. (OWZ).
Assigned age: mid Coniacian.
257
OTHER MACROFOSSILS
Among the various macrofossil groups, other thanammonites, only inoceramidbivalves seemtohave
the biostratigraphical potential to allow the establishment of a detailed separate zonal scheme. Even so, an
inoceramid zonation for Sergipe will probably only comprise the Turonian and Coniacian stages, since ino-
ceramids are not abundant enough in other parts of the sequence. Based on studies of a restricted geographic
area, Ressel (1988) recognized two distinct lower Turonian inoceramidassociations: a lower Mytiloides my-
tiloides andan upper Mytiloides hercynicus association. Todetermine whether theseassociations arelaterally
persistent and thus potential biozones requires further study. They appear tofall entirely within the ammonite
zone of Mammites nodosoides-Kamerunoceras turoniense. Acharacteristic andeasilyrecognizable taxon in
the lower inoceramid association is the new genus Rhyssomytiloides Ressel, 1988, which can then aid in the
identification of the nodosoides-turoniense Zone. Other inoceramid taxa described by Ressel (1988) do not
seem to be confmed to one zone only.
Kauffman & Bengtson (1985) listed a number of provisionally determined inoceramid taxa ; the
material is still under study and will eventually be integrated with the current biostratigraphical scheme.
Ofother macrofossil groups, only the echinoids have as yet received full systematic treatment (Smith
1991). Although a separate biozonation for echinoids is not justified, selected species were found to be
confined to one or two ammonite zones and thus potentially biostratigraphically useful (Smith & Bengtson
1991, fig. 3). For example, the occurrence of Micropedina olisiponensis (Forbes, 1850) in the lower upper
Cenomanian Pseudocalycoceras harpax-Thomelites aff. sornayi Zone is consistent with its occurrence in
Portugal and North Africa [Berthou &Lauverjat 1978 ; Smith & Bengtson 1988, fig. 3 (although wrongly
given as middle Cenomanian on pp. 11,32)].
PALAEOENVIRONMENTAL EVOLUTION
OF THE SERGIPE BASIN
Several authors have discussed the palaeoenvironmenta! development of the Sergipe Basin during the
Cretaceous, for example, Berthou & Bengtson (1988), Koutsoukos (1989), and Koutsoukos et al. (1991a,
1991b). We attempt here a synthesis of the time range that extends from the earliest marine incursion until
the end of the Cretaceous. Interpretations are based on our integratedbiostratigraphical work presented here,
within the currently available sedimentological and teetono-sedimentary framework.
The passage from the transitional, evaporitic phase to a fully marine drift phase was characterized by
progressive crusta! extension and thinning. This allowed for the establishment ofa narrowepicontinental sea,
coupled with the gradual development of oceanic conditions in the central South Atlantic. Open marine
conditions were established in Sergipe by late Aptian times, with neritic-oceanic circulation patterns, and
surface-water exchange between the central and South Atlantic oceans (Bengtson & Koutsoukos 1992),
possibly even at intermediate water-depths (epipelagic to mesopelagic) (Koutsoukos 1992). A mixed
carbonate-siliciclastic platform system (the Riachuelo Formation) developed from the late Aptian to late
Albian, with paralic (lagoonal and tidal flats) to upper bathyal environments. Intermittent dysoxic-anoxic
conditions occurred, with a maximum oxygen-depletion event in the late Aptian-earliest Albian (fig. 8). A
relative sea-level rise during the early Cenomanian caused the drowning of the shallow-water Riachuelo
platform. Fine-grained deep-water limestones of a carbonate ramp system (Cenomanian to middle Conia-
cian ; the Cotinguiba Formation), accumulated in neritic to upper bathyal environments, with moderately
dysoxic to truly anoxic bottom conditions and well oxygenated epipelagic water masses (Koutsoulcos et al.
1991b). Two maximum oxygen depletion events (dysaerobic to anaerobic) are recorded in the succession
(fig. 8) : in the early Cenomanian, and at the Cenomanian-Turonian boundary. More oxidized conditions
(dysaerobic to aerobic) are apparent from the middle-upper Albian and lower-middle Turonian deposits,
probably as aconsequence ofless restricted oceanic exchange between the central and South Atlantic oceans,
with the progressive establishment of open oceanic circulation patterns (Koutsoukos et al. 1991b).
In the late Coniacian-early Santonian a major oceanographic event ended the carbonate-dominated
depositional cycle in the Sergipe Basin. The final structural detachment of the South American and African
258
OTHER MACROFOSSILS
Among the various macrofossil groups, other thanammonites, only inoceramidbivalves seemtohave
the biostratigraphical potential to allow the establishment of a detailed separate zonal scheme. Even so, an
inoceramid zonation for Sergipe will probably only comprise the Turonian and Coniacian stages, since ino-
ceramids are not abundant enough in other parts of the sequence. Based on studies of a restricted geographic
area, Hessel (1988) recognized two distinct lower Turonian inoceramidassociations: a lower Mytiloides my-
tiloides andan upperMytiloides hercynicus association. Todetermine whether theseassociations arelaterally
persistent and thus potential biozones requires further study. They appear tofall entirely within the ammonite
zone of Mammites nodosoides-Kamerunoceras turoniense. Acharacteristic andeasilyrecognizable taxon in
the lower inoceramid association is the new genus Rhyssomytiioides Hessel, 1988, which can then aid in the
identification of the nodosoides-turoniense Zone. Other inoceramid taxa described by Hessel (1988) do not
seem to be confmed to one zone only.
Kauffman & Bengtson (1985) listed a number of provisionally determined inoceramid taxa ; the
material is still under study and will eventually be integrated with the current biostratigraphical scheme.
Ofother macrofossil groups, only the echinoids have as yet received full systematic treatment (Smith
1991). Although a separate biozonation for echinoids is not justified, selected species were found to be
confined to one or two ammonite zones and thus potentially biostratigraphically useful (Smith & Bengtson
1991, fig. 3). For example, the occurrence of Micropedina olisiponensis (Forbes, 1850) in the lower upper
Cenomanian Pseudocaiycoceras harpax-Thomelites aff. sornayi Zone is consistent with its occurrence in
Portugal and North Africa [Berthou & Lauverjat 1978 ; Smith & Bengtson 1988, fig. 3 (although wrongly
given as middle Cenomanian on pp. 11,32)].
PALAEOENVIRONMENTAL EVOLUTION
OF THE SERGIPE BASIN
Several authors have discussed the palaeoenvironmental development of the Sergipe Basin during the
Cretaceous, for example, Berthou & Bengtson (1988), Koutsoukos (1989), and Koutsoukos et ai. (1991a,
1991b). We attempt here a synthesis of the time range that extends from the earliest marine incursion until
the end of the Cretaceous. Interpretations are based on our integratedbiostratigraphical work presented here,
within the currently available sedimentological and tectono-sedimentary framework.
The passage from the transitional, evaporitic phase to a fully marine drift phase was characterized by
progressive crustal extension and thinning. This allowed for the establishment ofanarrowepicontinental sea,
coupled with the gradual development of oceanic conditions in the central South Atlantic. Open marine
conditions were established in Sergipe by late Aptian times, with neritic-oceanic circulation patterns, and
surface-water exchange between the central and South Atlantic oceans (Bengtson & Koutsoukos 1992),
possibly even at intermediate water-depths (epipelagic to mesopelagic) (Koutsoukos 1992). A mixed
carbonate-siliciclastic platform system (the Riachuelo Formation) developed from the late Aptian to late
Albian, with paralic (lagoonal and tidal flats) to upper bathyal environments. Intermittent dysoxic-anoxic
conditions occurred, with a maximum oxygen-depletion event in the late Aptian-earliest Albian (fig. 8). A
relative sea-level rise during the early Cenomanian caused the drowning of the shallow-water Riachuelo
platform. Fine-grained deep-water limestones of a carbonate ramp system (Cenomanian to middle Conia-
cian ; the Cotinguiba Formation), accumulated in neritic to upper bathyal environments, with moderately
dysoxic to truly anoxic bottom conditions and well oxygenated epipelagic water masses (Koutsoulcos et ai.
1991b). Two maximum oxygen depletion events (dysaerobic to anaerobic) are recorded in the succession
(fig. 8) : in the early Cenomanian, and at the Cenomanian-Turonian boundary. More oxidized conditions
(dysaerobic to aerobic) are apparent from the middle-upper Albian and lower-middle Turonian deposits,
probably as aconsequence ofless restricted oceanic exchange between the central and South Atlantic oceans,
with the progressive establishment of open oceanic circulation patterns (Koutsoukos et ai. 1991b).
In the late Coniacian-early Santonian a major oceanographic event ended the carbonate-dominated
depositional cycle in the Sergipe Basin. The final structural detachment of the South American and African
258
OTHER MACROFOSSILS
Among the various macrofossil groups, other thanammonites, only inoceramidbivalves seemtohave
the biostratigraphical potential to allow the establishment of a detailed separate zonal scheme. Even so, an
inoceramid zonation for Sergipe will probably only comprise the Turonian and Coniacian stages, since ino-
ceramids are not abundant enough in other parts of the sequence. Based on studies of a restricted geographic
area, Ressel (1988) recognized two distinct lower Turonian inoceramidassociations: a lower Mytiloides my-
tiloides andan upperMytiloides hercynicus association. Todetermine whether theseassociations arelaterally
persistent and thus potential biozones requires further study. They appear tofall entirely within the ammonite
zone of Mammites nodosoides-Kamerunoceras turoniense. Acharacteristic andeasilyrecognizable taxon in
the lower inoceramid association is the new genus Rhyssomytiloides Ressel, 1988, which can then aid in the
identification of the nodosoides-turoniense Zone. Other inoceramid taxa described by Ressel (1988) do not
seem to be confmed to one zone only.
Kauffman & Bengtson (1985) listed a number of provisionally determined inoceramid taxa ; the
material is still under study and will eventually be integrated with the current biostratigraphical scheme.
Ofother macrofossil groups, only the echinoids have as yet received full systematic treatment (Smith
1991). Although a separate biozonation for echinoids is not justified, selected species were found to be
confined to one or two ammonite zones and thus potentially biostratigraphically useful (Smith & Bengtson
1991, fig. 3). For example, the occurrence of Micropedina olisiponensis (Forbes, 1850) in the lower upper
Cenomanian Pseudocalycoceras harpax-Thomelites aff. sornayi Zone is consistent with its occurrence in
Portugal and North Africa [Berthou & Lauverjat 1978 ; Smith & Bengtson 1988, fig. 3 (although wrongly
given as middle Cenomanian on pp. 11,32)].
PALAEOENVIRONMENTAL EVOLUTION
OF THE SERGIPE BASIN
Several authors have discussed the palaeoenvironmenta! development of the Sergipe Basin during the
Cretaceous, for example, Berthou & Bengtson (1988), Koutsoukos (1989), and Koutsoukos et al. (1991a,
1991b). We attempt here a synthesis of the time range that extends from the earliest marine incursion until
the end of the Cretaceous. Interpretations are based on our integratedbiostratigraphical work presented here,
within the currently available sedimentological and teetono-sedimentary framework.
The passage from the transitional, evaporitic phase to a fully marine drift phase was characterized by
progressive crusta! extension and thinning. This allowed for the establishment ofanarrowepicontinental sea,
coupled with the gradual development of oceanic conditions in the central South Atlantic. Open marine
conditions were established in Sergipe by late Aptian times, with neritic-oceanic circulation patterns, and
surface-water exchange between the central and South Atlantic oceans (Bengtson & Koutsoukos 1992),
possibly even at intermediate water-depths (epipelagic to mesopelagic) (Koutsoukos 1992). A mixed
carbonate-siliciclastic platform system (the Riachuelo Formation) developed from the late Aptian to late
Albian, with paralic (lagoonal and tidal flats) to upper bathyal environments. Intermittent dysoxic-anoxic
conditions occurred, with a maximum oxygen-depletion event in the late Aptian-earliest Albian (fig. 8). A
relative sea-level rise during the early Cenomanian caused the drowning of the shallow-water Riachuelo
platform. Fine-grained deep-water limestones of a carbonate ramp system (Cenomanian to middle Conia-
cian ; the Cotinguiba Formation), accumulated in neritic to upper bathyal environments, with moderately
dysoxic to truly anoxic bottom conditions and well oxygenated epipelagic water masses (Koutsoulcos et al.
1991b). Two maximum oxygen depletion events (dysaerobic to anaerobic) are recorded in the succession
(fig. 8) : in the early Cenomanian, and at the Cenomanian-Turonian boundary. More oxidized conditions
(dysaerobic to aerobic) are apparent from the middle-upper Albian and lower-middle Turonian deposits,
probably as aconsequence ofless restricted oceanic exchange between the central and South Atlantic oceans,
with the progressive establishment of open oceanic circulation patterns (Koutsoukos et al. 1991b).
In the late Coniacian-early Santonian a major oceanographic event ended the carbonate-dominated
depositional cycle in the Sergipe Basin. The final structural detachment of the South American and African
258
plates restrltMin theestablishment ofadeep-ocean circulationregime. The basinwas abruptly tilted seawards
along a NE-SW-trending rotational axis, which caused uplift of the north-western margin of the basin and
created a prominent submarine topography (Koutsoukos & Hart 1990, fig. 13). Simultaneously there was a
rapid change from a dominantly dry to a more humid and warmclimate, probably causedby the development
of a high-pressure atmospheric cell at low latitudes over the widening Atlantic watermasses (e.g., Parrish &
Curtis 1982, Chang et al. 1988). This climatic turnover caused increased siliciclastic influx to the basin (due
to continental runoft), the material of which wasderived from the upliftedareas in the north-west.
Thick successions of shales and, suboidimltely, turbiditic sandstones weredeposited; these form theCalumbi.
Member of the Formation.
Stages
Maastrichtian
Campanian
Santonian
Coniacian
Turonian
Cenomanian
Albian
Upper Aptian
Planktonic
foraminiferal
zones
C. contusa-G. aegyptiaca
G. gansseri-G. stuartiformis
C. ex gr. fornicata-
G. linneiana
.G. orientalis-G. ventricosa
G. patelliformis-G. elevatal
stuartifonnis plexus
Dicarinella asymetrica
D concavata-M sinuosa
A. cretacea-D. primitiva
Dicarinella primitiva
H. aprica-
H. (W) baltica
H. (W) archaeocretacea-
H. reussi
H. (W) aprJca-G. bentonensis
H(W) baltica-
H. (W) brittonensis
P delrioensis-R. appennil1ica
R. brotzeni
H. (H.) gorbachlkae-
T raynaudi
G. texomaensis-
B. breggiensis
B. breggiel1sis-
.' T ex gr. primula
T ex gr. prJmula-
T bejaouaensis
T beJaouaensis
G. cushmani-
T bejaouaensis
G. ex gr. maridalensls-H (H.) slmilis
G. barri-H. (H.] gorbachlkae
NW
Palaeobathymetric
evolution
SE
,
,
,
: :: :.: :.:.:.: :.;.: :.:.: :::::::.
I
>,'
cnlQl
"0, c
EI'&
35 'I <5
,"55
0.1
Fig. 8 - Palaeobathymetric evolution of the Serglpe Basin. Shaded areas Indicate time of maximumdysoxlc-anoxic
episodes. The left-hand side of the diagram corresponds to the present-day, north-western onshore border of the basin
(Riachuelo and ltaporanga highs), and the right-hand side to the south-eastern limit of the studied area (off-shore area of
welll-SES-3 : Koutsoukos 1992, fig. 1) (from Koutspukos 1992, fig. 2(E) ; stage thicknesses not to scale). '
259
plates restrltMin theestablishment ofadeep-ocean circulationregime. The basinwas abruptly tilted seawards
along a NE-SW-trending rotational axis, which caused uplift of the north-western margin of the basin and
created a prominent submarine topography (Koutsoukos & Hart 1990, fig. 13). Simultaneously there was a
rapid change from a dominantly dry to a more humid and warmclimate, probably causedby the development
of a high-pressure atmospheric cell at low latitudes over the widening Atlantic watermasses (e.g., Parrish &
Curtis 1982, Chang et al. 1988). This climatic turnover caused increased siliciclastic influx to the basin (due
to intensificontinental runoft), the material of which wasderived from the upliftedareas in the north-west.
Thick successionsof shales and, suboidimltely, turbiditic sandstones weredeposited; these form theCalumbi.
Member of the P i ~ a b u ~ u Formation.
Stages
Maastrichtian
Campanian
Santonian
Coniacian
Turonian
Cenomanian
Albian
Upper Aptian
Planktonic
foraminiferal
zones
C. contusa-G. aegyptiaca
G. gansseri-G. stuartiformis
C. ex gr. fornicata-
G. linneiana
.G. orientalis-G. ventricosa
G. patelliformis-G. elevata!
stuartifonnis plexus
Oicarinella asymetrica
o concavata-M sinuosa
A. cretacea-O. primitiva
Oicarinella primitiva
H. aprica-
H. (W) baltica
H. (W) archaeocretacea-
H. reussi
H. (W) aprJca-G. bentonensis
H(W) baftica-
H. (W) brittonensis
P delrioensis-R. appennil1ica
R. brotzeni
H. (H.) gorbachlkae-
T raynaudi
G. texomaensis-
B. breggiensis
B. breggiel1sis-
.' T ex gr. primula
T ex gr. prJmula-
T bejaouaensis
T beJaouaensis
G. cushmani-
T bejaouaensis
G. ex gr. maridalensls-H (H.i slmilis
G. barri-H. (H.] gorbachlkae
NW
Palaeobathymetric
evolution
SE
,
,
,
: :: :.: :.:.:.: :.;.: :.:.: :::::::.
I
>,'
(\jIm
"0, c
EI'&
35 'I <5
~ ,"55
0.1
Fig. 8 - Palaeobathymetric evolution of the Serglpe Basin. Shaded areas Indicate time of maximumdysoxlc-anoxic
episodes. The left-hand side of the diagram corresponds to the present-day, north-western onshore border of the basin
(Riachuelo and ltaporanga highs), and the right-hand side to the south-eastern limit of the studied area (off-shore area of
welll-SES-3 : Koutsoukos 1992, fig. 1) (from Koutspukos 1992, fig. 2(E) ; stage thicknesses not to scale). '
259
plates restrltMin theestablishment ofadeep-ocean circulationregime. The basinwas abruptly tilted seawards
along a NE-SW-trending rotational axis, which caused uplift of the north-western margin of the basin and
created a prominent submarine topography (Koutsoukos & Hart 1990, fig. 13). Simultaneously there was a
rapid change from a dominantly dry to a more humid and warmclimate, probably causedby the development
of a high-pressure atmospheric cell at low latitudes over the widening Atlantic watermasses (e.g., Parrish &
Curtis 1982, Chang et al. 1988). This climatic turnover caused increased siliciclastic influx to the basin (due
to continental runoft), the material of which wasderived from the upliftedareas in the north-west.
Thick successions of shales and, suboidimltely, turbiditic sandstones weredeposited; these form theCalumbi.
Member of the Formation.
Stages
Maastrichtian
Campanian
Santonian
Coniacian
Turonian
Cenomanian
Albian
Upper Aptian
Planktonic
foraminiferal
zones
C. contusa-G. aegyptiaca
G. gansseri-G. stuartiformis
C. ex gr. fornicata-
G. linneiana
.G. orientalis-G. ventricosa
G. patelliformis-G. elevatal
stuartifonnis plexus
Dicarinella asymetrica
D concavata-M sinuosa
A. cretacea-D. primitiva
Dicarinella primitiva
H. aprica-
H. (W) baltica
H. (W) archaeocretacea-
H. reussi
H. (W) aprJca-G. bentonensis
H(W) baltica-
H. (W) brittonensis
P delrioensis-R. appennil1ica
R. brotzeni
H. (H.) gorbachlkae-
T raynaudi
G. texomaensis-
B. breggiensis
B. breggiel1sis-
.' T ex gr. primula
T ex gr. prJmula-
T bejaouaensis
T beJaouaensis
G. cushmani-
T bejaouaensis
G. ex gr. maridalensls-H (H.) slmilis
G. barri-H. (H.] gorbachlkae
NW
Palaeobathymetric
evolution
SE
,
,
,
: :: :.: :.:.:.: :.;.: :.:.: :::::::.
I
>,'
cnlQl
"0, c
EI'&
35 'I <5
,"55
0.1
Fig. 8 - Palaeobathymetric evolution of the Serglpe Basin. Shaded areas Indicate time of maximumdysoxlc-anoxic
episodes. The left-hand side of the diagram corresponds to the present-day, north-western onshore border of the basin
(Riachuelo and ltaporanga highs), and the right-hand side to the south-eastern limit of the studied area (off-shore area of
welll-SES-3 : Koutsoukos 1992, fig. 1) (from Koutspukos 1992, fig. 2(E) ; stage thicknesses not to scale). '
259
Apalaeobathymebic maximumappears to haveoccurred in the early Campanian (fig. 8) (Koutsoukos
1989), when the bathymebically deepest environments are recorded in all the sites studied. Foraminiferal
speciesdiversityreachedapeakin theCampanian, probably in responseto increasednicheavailability, which
promoted evolutionary diversification.
Palaeoenvironmental conclusions
The marine Cretaceous palaeoenvironmental evolutionofthe basincanbesubdividedintothree major
phases: (1) a late Aptian to late Albian paralic to upper bathyal phase (mixed carbonate-siliciclastic platform
system; Riachuelo Formation) ; (2) an early Cenomanian to mid-Coniacian neritic to upper bathyal phase
(carbonate ramp system; Cotinguiba Formation) ; and (3) a late Coniacian or Santonian to late Maasbichtian
middle-deep neritic to lower bathyal phase(siliciclastic system; Calumbi Member Formation).
Foraminiferal species diversity reachedapeak in the Campanian. This was associated with a palaeobathyme-
bic maximum, probably in the early Campanian, from when the deepest environments are recordedat all the
sites studied.
Acknowledgements
We thank Petr61eo Brasileiro S.A. (petrobnis), Rio de Janeiro, for permission to publish the paper.
Paleontologiska institutionen, Uppsala University (Sweden), provided working facilities for P. B. during
research visits in 1991-1992. Marta C. Viviers (petrobnis) advised on the microfossil zonations, Suzana I.
Bengtson (Uppsala) revised determinations of the lower Cenomanian ammonites, and Pierre-Yves Berthou
(Paris) contributedstratigraphical data onechinoids. We thankP.Y. BerthouandFrancisRobaszynski (Mons)
for reviewing the manuscript for publication. The final drafting of the illustrations was by Rolf Koch and
Rainer Ziihlke (Universitat Heidelberg), using Aldus FreeHand on a Macintosh Quadra 950. Part of the
underlying work was financed by grants to P. B. from the SwedishNatural ScienceResearch Council (NFR).
REFERENCES CITED
ASMUS H. E. &BAISCH P. R. (1983) - Geological evolution of the Brazilian continental margin. Episodes, 4,
p.3-9.
BANDEIRA A. N., Jr. (1978) - Sedimentologia e microfacies calcarias das Riachuelo e Cotinguiba
da Bacia Sergipe/Alagoas. Bol. Tee. Petrobras, 21, I, p. 17-69.
BENGrSON P. (1979) - Abioestratigrafia esquecida : dos metodos bioestratigraficos no Cretaceo
medio do BrasH. Anais Acad. Brasil. Cienc., 51, 3, p. 535-544.
BENGrSON P. (1983) - The Cenomanian-Coniacian of the Sergipe Basin, Brazil. Fossils and Strata, 12, p. 1-
78.
BENGrSON P. & KOursOUKOS E. A. M. (1992) - Ammonite and foraminiferal dating of the fIrst marine con-
nection between the central and South Atlantic. In R. Curnelle (ed.), Geologie Africaine, Compte-
rendu des Colloques de geologie de Libreville, 68 Mai, 1991. Bull. Centr. Rech. Explor.- Produc. Elf
Aquitaine. Mem. 13, p. 403.
BmmmuP.-Y. (1984) - Albian-Turonian stageboundaries and subdivisions in the western Portuguese Basin,
with special emphasis on theCenomanian-Turonianboundaryin the ammonitefacies andrudist facies.
Bull. Geol. Soc. Denmark, 33,1-2, p. 41-55.
BBRTHOUP.-Y. &BENGrSONP. (1988) - Slratigraphic correlationbymicrofacies ofthe Cenomanian-Coniacian
of the Sergipe Basin, Brazil. Fossils and Strata, 21, p. 1-88.
BBRTIIouP.-Y. &LAUVERJATJ. (1978) -Lebassinoccidental portugaisde I'AlbienauCampanien.AnnalsMus.
Hist. Nat. Nice, 4 [for 1976], p. 1.1-1.14.
BEURLENG. (1969) - Afauna do complexo Riachuelo/Maruim : I - Ammonoidea. Bol. Tee. Petrobras, 11, 4
[for 1968], p. 437-482.
BEURLEN. G. (1970) - Uma nova fauna de amon6ides da SapucarilLaranjeiras (Cret8ceo de Ser-
gipe) : sabre sua bioesttatigrafia. Bol. Tee. Petrobras, 12,2 [for 1969], p.147-169.
BEURLEN K. (1961) - Die Kreide im KUstenbereich von Sergipe bis Paraiba do Norte (Brasilien). Zeitschr.
Deutsch. Geol. Gesells., 112,3 [for 1960], p. 378-384.
260
Apalaeobathymetric maximumappears to haveoccurred in the early Campanian (fig. 8) (Koutsoukos
1989), when the bathymetrically deepest environments are recorded in all the sites studied. Foraminiferal
speciesdiversityreachedapeakin theCampanian, probably in responseto increasednicheavailability, which
promoted evolutionary diversification.
Palaeoenvironmental conclusions
The marine Cretaceous palaeoenvironmental evolutionofthe basincanbesubdividedintothree major
phases: (1) a late Aptian to late Albian paralic to upper bathyal phase (mixed carbonate-siliciclastic platform
system; Riachuelo Formation) ; (2) an early Cenomanian to mid-Coniacian neritic to upper bathyal phase
(carbonate ramp system; Cotinguiba Formation) ; and (3) a late Coniacian or Santonian to late Maastrichtian
middle-deep neritic to lower bathyal phase (siliciclastic system; Calumbi Member Formation).
Foraminiferal species diversity reachedapeak in the Campanian. This was associated with a palaeobathyme-
tric maximum, probably in the early Campanian, from when the deepest environments are recordedat all the
sites studied.
Acknowledgements
We thank Petr61eo Brasileiro S.A. (petrobnis), Rio de Janeiro, for permission to publish the paper.
Paleontologiska institutionen, Uppsala University (Sweden), provided working facilities for P. B. during
research visits in 1991-1992. Marta C. Viviers (petrobras) advised on the microfossil zonations, Suzana I.
Bengtson (Uppsala) revised determinations of the lower Cenomanian ammonites, and Pierre-Yves Berthou
(Paris) contributedstratigraphical data onechinoids. We thankP.Y. BerthouandFrancisRobaszynski (Mons)
for reviewing the manuscript for publication. The final drafting of the illustrations was by Rolf Koch and
Rainer ZUhlke (Universitat Heidelberg), using Aldus FreeHand on a Macintosh Quadra 950. Part of the
underlying work was financed by grants to P. B. from the SwedishNatural ScienceResearch Council (NFR).
REFERENCES CITED
ASMUS H. E. &BAISCH P. R. (1983) - Geological evolution of the Brazilian continental margin. Episodes, 4,
p.3-9.
BANDEIRA A. N., Jr. (1978) - Sedimentologia e microfacies calcarias das Riachuelo e Cotinguiba
da Bacia Sergipe/Alagoas. Bol. Tee. Petrobras, 21, I, p. 17-69.
BENGrSON P. (1979) - Abioestratigrafia esquecida : dos metodos bioestratigraficos no Cretaceo
medio do Brasil. Anais Acad. Brasil. Cienc., 51, 3, p. 535-544.
BENGrSON P. (1983) - The Cenomanian-Coniacian of the Sergipe Basin, Brazil. Fossils and Strata, 12, p. 1-
78.
BENGrSON P. & KOursOUKOS E. A. M. (1992) - Ammonite and foraminiferal dating of the fIrst marine con-
nection between the central and South Atlantic. In R. Curnelle (ed.), Geologie Africaine, Compte-
rendu des Colloques de geologie de Libreville, 68 Mai, 1991. Bull. Centro Rech. Explor.- Produc. Elf
Aquitaine. Mem. 13, p. 403.
BmmmuP.-Y. (1984) - Albian-Turonian stageboundaries and subdivisions in the western Portuguese Basin,
with special emphasis on theCenomanian-Turonianboundaryin the ammonitefacies andrudist facies.
Bull. Geol. Soc. Denmark, 33,1-2, p. 41-55.
BBRTHOUP.-Y. &BENGrSONP. (1988) - Stratigraphic correlationbymicrofacies ofthe Cenomanian-Coniacian
of the Sergipe Basin, Brazil. Fossils and Strata, 21, p. 1-88.
BBRTIIouP.-Y. &LAUVERJATJ. (1978) -Lebassinoccidental portugais de I'AlbienauCampanien.AnnalsMus.
Hist. Nat. Nice, 4 [for 1976], p. 1.1-1.14.
BEURLENG. (1969) - Afauna do complexo Riachuelo/Maruim : I - Ammonoidea. Bol. Tee. Petrobras, II, 4
[for 1968], p. 437-482.
BEURLEN. G. (1970) - Uma nova fauna de amon6ides da SapucarilLaranjeiras (Cret8ceo de Ser-
gipe) : sObre sua bioestratigrafia. Bol. Tee. Petrobras, 12,2 [for 1969], p.147-169.
BEURLEN K. (1961) - Die Kreide im KUstenbereich von Sergipe bis Paraiba do Norte (Brasilien). Zeitschr.
Deutsch. Geol. Gesells., 112,3 [for 1960], p. 378-384.
260
Apalaeobathymebic maximumappears to haveoccurred in the early Campanian (fig. 8) (Koutsoukos
1989), when the bathymebically deepest environments are recorded in all the sites studied. Foraminiferal
speciesdiversityreachedapeakin theCampanian, probably in responseto increasednicheavailability, which
promoted evolutionary diversification.
Palaeoenvironmental conclusions
The marine Cretaceous palaeoenvironmental evolutionofthe basincanbesubdividedintothree major
phases: (1) a late Aptian to late Albian paralic to upper bathyal phase (mixed carbonate-siliciclastic platform
system; Riachuelo Formation) ; (2) an early Cenomanian to mid-Coniacian neritic to upper bathyal phase
(carbonate ramp system; Cotinguiba Formation) ; and (3) a late Coniacian or Santonian to late Maasbichtian
middle-deep neritic to lower bathyal phase (siliciclastic system; Calumbi Member Formation).
Foraminiferal species diversity reachedapeak in the Campanian. This was associated with a palaeobathyme-
bic maximum, probably in the early Campanian, from when the deepest environments are recordedat all the
sites studied.
Acknowledgements
We thank Petr61eo Brasileiro S.A. (petrobnis), Rio de Janeiro, for permission to publish the paper.
Paleontologiska institutionen, Uppsala University (Sweden), provided working facilities for P. B. during
research visits in 1991-1992. Marta C. Viviers (petrobras) advised on the microfossil zonations, Suzana I.
Bengtson (Uppsala) revised determinations of the lower Cenomanian ammonites, and Pierre-Yves Berthou
(Paris) contributedstratigraphical data onechinoids. We thankP.Y. BerthouandFrancisRobaszynski (Mons)
for reviewing the manuscript for publication. The final drafting of the illustrations was by Rolf Koch and
Rainer Ziihlke (Universitat Heidelberg), using Aldus FreeHand on a Macintosh Quadra 950. Part of the
underlying work was financed by grants to P. B. from the SwedishNatural ScienceResearch Council (NFR).
REFERENCES CITED
ASMUS H. E. &BAISCH P. R. (1983) - Geological evolution of the Brazilian continental margin. Episodes, 4,
p.3-9.
BANDEIRA A. N., Jr. (1978) - Sedimentologia e microfacies calcarias das Riachuelo e Cotinguiba
da Bacia Sergipe/Alagoas. Bol. Tee. Petrobras, 21, I, p. 17-69.
BENGrSON P. (1979) - Abioestratigrafia esquecida : dos metodos bioestratigraficos no Cretaceo
medio do BrasH. Anais Acad. Brasil. Cienc., 51, 3, p. 535-544.
BENGrSON P. (1983) - The Cenomanian-Coniacian of the Sergipe Basin, Brazil. Fossils and Strata, 12, p. 1-
78.
BENGrSON P. & KOursOUKOS E. A. M. (1992) - Ammonite and foraminiferal dating of the fIrst marine con-
nection between the central and South Atlantic. In R. Curnelle (ed.), Geologie Africaine, Compte-
rendu des Colloques de geologie de Libreville, 68 Mai, 1991. Bull. Centr. Rech. Explor.- Produc. Elf
Aquitaine. Mem. 13, p. 403.
BmmmuP.-Y. (1984) - Albian-Turonian stageboundaries and subdivisions in the western Portuguese Basin,
with special emphasis on theCenomanian-Turonianboundaryin the ammonitefacies andrudist facies.
Bull. Geol. Soc. Denmark, 33,1-2, p. 41-55.
BBRTHOUP.-Y. &BENGrSONP. (1988) - Slratigraphic correlationbymicrofacies ofthe Cenomanian-Coniacian
of the Sergipe Basin, Brazil. Fossils and Strata, 21, p. 1-88.
BBRTIIouP.-Y. &LAUVERJATJ. (1978) -Lebassinoccidental portugaisde I'AlbienauCampanien.AnnalsMus.
Hist. Nat. Nice, 4 [for 1976], p. 1.1-1.14.
BEURLENG. (1969) - Afauna do complexo Riachuelo/Maruim : I - Ammonoidea. Bol. Tee. Petrobras, 11, 4
[for 1968], p. 437-482.
BEURLEN. G. (1970) - Uma nova fauna de amon6ides da SapucarilLaranjeiras (Cret8ceo de Ser-
gipe) : sabre sua bioestratigrafia. Bol. Tee. Petrobras, 12,2 [for 1969], p.147-169.
BEURLEN K. (1961) - Die Kreide im KUstenbereich von Sergipe bis Paraiba do Norte (Brasilien). Zeitschr.
Deutsch. Geol. Gesells., 112,3 [for 1960], p. 378-384.
260
CARON M. (1985) - Cretaceous planktic foraminifera.ln BoLU H. M., SAUNDERS J. B. & PERcH-NIELSEN K.
(eds), Plankton Stratigraphy, p. 17-86. Cambridge University Press, London.
CHANG H. K., KOWSMANN R. O. & FIGUEIREDO A. M. F. DE (1988) - Newconcepts on the development of east
Brazilian marginal basins. Episodes, 11,3, p. 194-202.
FEu6 F. J. (1980) - Estudo dos carbonatos Muribeca e Riachuelo no Alto de Aracaju - Bacia Sergipe/Alagoas
- nordeste do Brasil. Anais XXXI Congr. Brasil. Geol. (Camboriu. SC). 1, p. 320-332.
FRErrAS L. C. S. (1984) - Nanof6sseis calc8rios e sua (Aptiano-Mioceno) na Bacia Sergipe/
Alagoas. Unpublished M. Sc. dissertation. Departarnento de Geociencias da Universidade Federal do
Rio de Janeiro, Rio de Janeiro.
HANCOCK J. M. (1991) - Ammonite scales for the Cretaceous System. Cret. Res., 12,3, p. 259-291.
HEDBERG H. D. (1976) - International Stratigraphic Guide: a guide to Stratigraphic Classification, Termino-
logy and Procedure. 200 pp., John Wiley & Sons, New York.
HESSEL M. H. R. (1988) - Lower Turonian inoceramids from Sergipe, Brazil: systematics, stratigraphy and
palaeoecology. Fossils and Strata, 22, p. 1-49.
KAUFFMANE. G: & BENGTSON P. (1985) - Mid-Cretaceous inoceramids from Sergipe, Brazil: a progress report.
Cret. Res., 6, 3, p. 311-315.
KENNEDY W. J. & COBBAN W. A. (1991) - Stratigraphy and interregional correlation of the Cenomanian-
Turonian transition in the Western Interior of the United States near Pueblo, Colorado, a potential
boundary stratotype for the base of the Turonian Stage. Newsl. Strat., 24, 1/2, p. 1-33.
KoursoUKOS E. A. M. (1989) - Mid- to late Cretaceous microbiostratigraphy, palaeo-ecology and palaeogeo-
graphy of the Sergipe Basin, northeastern Brazil. Unpublished Ph.D. thesis. 2 vols, 886 pp., Council
for National Academic Awards/polytechnic South West, Plymouth.
KOurSOUKOS E. A. M. (1992) - Late Aptian to Maastrichtian foraminiferal biogeography andpalaeoceanogra-
phy of the Sergipe Basin, Brazil. Palaeogeogr., Palaeoclim., Palaeoecol., 92, p. 295-324.
KOursoUKOS E. A. M., DESTRO N., AzAMBUJA FILHO N. C. DE & SPADINI A. R. (in press) - Upper Aptian-Iower
Coniacian carbonate sequences in the Sergipe Basin, northeastern Brazil. In T. SIMO, B. W. SCOIT &
J.-P. MASSE (eds), Cretaceous Carbonate Platforms. Mem. Amer. Ass. Petrol. Geol.
KOurSOUKOS E. A. M. & HART M. B. (1990) - Cretaceous foraminiferal morphogroup distribution patterns,
palaeocommunities and trophic structures: a case study from the Sergipe Basin, Brazil. Trans. R. Soc.
Edinburgh, Earth Sci., 81, p. 221-246.
KOurSOUKOS E. A. M., MELLO M. R., AzAMBUJAFILHON. C. DE, HARTM. B. & MAxWELLJ. R. (1991a) - The
upper Aptian-Albian succession of the Sergipe Basin, Brazil : an integrated paleoenvironmental
assessment Amer. Ass. Petrol. Geol. Bull., 75, 3, p. 479-498.
KOurSOUKOS E. A. M., MELLO M. R. & AzAMBUJA FILHO N. C. DE (1991b) - Micropalaeontological and geo-
chemical evidence of mid-Cretaceous hypoxic-anoxic environments in the Sergipe Basin, northeas-
tern Brazil. I n R. V. TYSON & T. H. PEARSON (eds), Modern and ancient continental shelfanoxia. Geol.
Soc. Spec. Publ., 58, p. 427-447.
LANA M. C. (1990) - Bacia de Sergipe-Alagoas: uma hip6tese de tectono-sedirnentar.ln G. P. R.
GABAGLIA & E. J. M!LANr (eds), Origem e de bacias sedimentares, p. 311-332. Ed. Gavea:
R. Redisch Prog. Visual Prod. Graf. e : Petrobras, Rio de Janeiro.
MASTERS B. A. (1977) - Mesowic planktonic Foraminifera: a world-wide review and analysis. In RAMSAY A.
T. S. (ed.), Oceanic Micropal., I, p. 301-731. Academic Press, London.
MAURY C. J. (1937) - 0 Cretaceo de Sergipe. Serv. Geol. Miner. Brasil, Monogr., 11, p. 1-283.
OJEDA H. A. O. (1982) - Structural framework, stratigraphy and evolution of Brazilian marginal basins. Bull.
Amer. Ass. Petrol. Geol., 66, 6, p. 732-749.
OJEDA H.A. O. & FUGrrAA. M. (1976) - BaciaSergipe/Alagoas : Geologiaregional e perspectivas petroliferas.
Anais XXVIII Congr. (Brasil. Geol.) [porto Alegre, RS], I, p. 137-158.
PARRISH J. T. & CURTIS R. L. (1982) - Atmospheric circulation, upwelling, and organic-rich rocks in the
Mesozoic and Cenowic Eras. Palaeogeogr., Palaeoclim., Palaeoecol., 40,1-3: p. 31-66..
PETRI S. (1962) - Foraminiferos cretaceos de Sergipe. Bol. Fac. Filos., Cienc. Letras, Univ. Sa.o Paulo, 265
(Geologia, 20), p. 1-140.
PONTE F. C. & ASMUS H. E. (1976) - The Brazilian marginal basins: current state of knowledge. In ALMEIDA
F. F. M. DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc., 48, Suplem.,p. 215-
239.
261
CARON M. (1985) - Cretaceous planktic foraminifera. In BoLU H. M., SAUNDERS J. B. & PERCH-NIELSEN K.
(eds), Plankton Stratigraphy, p. 17-86. Cambridge University Press, London.
CHANG H. K., KOWSMANN R. O. & FIGUEIREDO A. M. F. DE (1988) - Newconcepts on the development of east
Brazilian marginal basins. Episodes, 11,3, p. 194-202.
FEu6 F. J. (1980) - Estudo dos carbonatos Muribeca e Riachuelo no Alto de Aracaju - Bacia Sergipe/Alagoas
- nordeste do Brasil. Anais XXXI Congr. Brasil. Geol. (Camboriu. SC). 1, p. 320-332.
FRErrAS L. C. S. (1984) - Nanof6sseis calc8rios e sua (Aptiano-Mioceno) na Bacia Sergipe/
Alagoas. Unpublished M. Sc. dissertation. Departamento de Geociencias da Universidade Federal do
Rio de Janeiro, Rio de Janeiro.
HANCOCK J. M. (1991) - Ammonite scales for the Cretaceous System. Cret. Res., 12,3, p. 259-291.
HEDBERG H. D. (1976) - International Stratigraphic Guide: a guide to Stratigraphic Classification, Termino-
logy and Procedure. 200 pp., John Wiley & Sons, New York.
HESSEL M. H. R. (1988) - Lower Turonian inoceramids from Sergipe, Brazil: systematics, stratigraphy and
palaeoecology. Fossils and Strata, 22, p. 1-49.
KAUFFMANE. G: & BENGTSON P. (1985) - Mid-Cretaceous inoceramids from Sergipe, Brazil: a progress report.
Cret. Res., 6, 3, p. 311-315.
KENNEDY W. J. & COBBAN W. A. (1991) - Stratigraphy and interregional correlation of the Cenomanian-
Turonian transition in the Western Interior of the United States near Pueblo, Colorado, a potential
boundary stratotype for the base of the Turonian Stage. Newsl. Strat., 24, 1/2, p. 1-33.
KoursoUKOS E. A. M. (1989) - Mid- to late Cretaceous microbiostratigraphy, palaeo-ecology and palaeogeo-
graphy of the Sergipe Basin, northeastern Brazil. Unpublished Ph.D. thesis. 2 vols, 886 pp., Council
for National Academic Awards/polytechnic South West, Plymouth.
KOurSOUKOS E. A. M. (1992) - Late Aptian to Maastrichtian foraminiferal biogeography andpalaeoceanogra-
phy of the Sergipe Basin, Brazil. Palaeogeogr., Palaeoclim., Palaeoecol., 92, p. 295-324.
KOursoUKOS E. A. M., DESTRO N., AzAMBUJA FILHO N. C. DE & SPADINI A. R. (in press) - Upper Aptian-lower
Coniacian carbonate sequences in the Sergipe Basin, northeastern Brazil. In T. SIMO, B. W. SCOTT &
J.-P. MASSE (eds), Cretaceous Carbonate Platforms. Mem. Amer. Ass. Petrol. Geol.
KOurSOUKOS E. A. M. & HART M. B. (1990) - Cretaceous foraminiferal morphogroup distribution patterns,
palaeocommunities and trophic structures: a case study from the Sergipe Basin, Brazil. Trans. R. Soc.
Edinburgh, Earth Sci., 81, p. 221-246.
KOurSOUKOS E. A. M., MELLO M. R., AzAMBUJAFILHON. C. DE, HARTM. B. & MAxWELLJ. R. (1991a) - The
upper Aptian-Albian succession of the Sergipe Basin, Brazil : an integrated paleoenvironmental
assessment Amer. Ass. Petrol. Geol. Bull., 75, 3, p. 479-498.
KOurSOUKOS E. A. M., MELLO M. R. & AzAMBUJA FILHO N. C. DE (1991b) - Micropalaeontological and geo-
chemical evidence of mid-Cretaceous hypoxic-anoxic environments in the Sergipe Basin, northeas-
tern Brazil.lnR. V. TYSON & T. H. PEARSON (eds), Modern and ancient continental shelfanoxia. Geol.
Soc. Spec. Publ., 58, p. 427-447.
LANA M. C. (1990) - Bacia de Sergipe-Alagoas: uma hip6tese de tectono-sedirnentar.ln G. P. R.
GABAGLIA & E. J. M!LANr (eds), Origem e de bacias sedimentares, p. 311-332. Ed. Gavea:
R. Redisch Prog. Visual Prod. Graf. e : Petrobras, Rio de Janeiro.
MASTERS B. A. (1977) - Mesowic planktonic Foraminifera: a world-wide review and analysis. In RAMSAY A.
T. S. (ed.), Oceanic Micropal., I, p. 301-731. Academic Press, London.
MAURY C. J. (1937) - 0 Cretaceo de Sergipe. Servo Geol. Miner. Brasil, Monogr., 11, p. 1-283.
OJEDA H. A. O. (1982) - Structural framework, stratigraphy and evolution of Brazilian marginal basins. Bull.
Amer. Ass. Petrol. Geol., 66, 6, p. 732-749.
OJEDA H..A. O. & FUGrrAA. M. (1976) - BaciaSergipe/Alagoas : Geologiaregional e perspectivas petroliferas.
Anais XXVIII Congr. (Brasil. Geol.) [porto Alegre, RS], I, p. 137-158.
PARRISH J. T. & CURTIS R. L. (1982) - Atmospheric circulation, upwelling, and organic-rich rocks in the
Mesozoic and Cenowic Eras. Palaeogeogr., Palaeoclim., Palaeoecol., 40,1-3: p. 31-66..
PETRI S. (1962) - Foraminiferos cretaceos de Sergipe. Bol. Fac. Filos., Cienc. Letras, Univ. Sa.o Paulo, 265
(Geologia, 20), p. 1-140.
PONTE F. C. & ASMUS H. E. (1976) - The Brazilian marginal basins: current state of knowledge. In ALMEIDA
F. F. M. DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc., 48, Suplem.,p. 215-
239.
261
CARON M. (1985) - Cretaceous planktic foraminifera.ln BoLU H. M., SAUNDERS J. B. & PERcH-NIELSEN K.
(eds), Plankton Stratigraphy, p. 17-86. Cambridge University Press, London.
CHANG H. K., KOWSMANN R. O. & FIGUEIREDO A. M. F. DE (1988) - Newconcepts on the development of east
Brazilian marginal basins. Episodes, 11,3, p. 194-202.
FEu6 F. J. (1980) - Estudo dos carbonatos Muribeca e Riachuelo no Alto de Aracaju - Bacia Sergipe/Alagoas
- nordeste do Brasil. Anais XXXI Congr. Brasil. Geol. (Camboriu. SC). 1, p. 320-332.
FRErrAS L. C. S. (1984) - Nanof6sseis calc8rios e sua (Aptiano-Mioceno) na Bacia Sergipe/
Alagoas. Unpublished M. Sc. dissertation. Departarnento de Geociencias da Universidade Federal do
Rio de Janeiro, Rio de Janeiro.
HANCOCK J. M. (1991) - Ammonite scales for the Cretaceous System. Cret. Res., 12,3, p. 259-291.
HEDBERG H. D. (1976) - International Stratigraphic Guide: a guide to Stratigraphic Classification, Termino-
logy and Procedure. 200 pp., John Wiley & Sons, New York.
HESSEL M. H. R. (1988) - Lower Turonian inoceramids from Sergipe, Brazil: systematics, stratigraphy and
palaeoecology. Fossils and Strata, 22, p. 1-49.
KAUFFMANE. G: & BENGTSON P. (1985) - Mid-Cretaceous inoceramids from Sergipe, Brazil: a progress report.
Cret. Res., 6, 3, p. 311-315.
KENNEDY W. J. & COBBAN W. A. (1991) - Stratigraphy and interregional correlation of the Cenomanian-
Turonian transition in the Western Interior of the United States near Pueblo, Colorado, a potential
boundary stratotype for the base of the Turonian Stage. Newsl. Strat., 24, 1/2, p. 1-33.
KoursoUKOS E. A. M. (1989) - Mid- to late Cretaceous microbiostratigraphy, palaeo-ecology and palaeogeo-
graphy of the Sergipe Basin, northeastern Brazil. Unpublished Ph.D. thesis. 2 vols, 886 pp., Council
for National Academic Awards/polytechnic South West, Plymouth.
KOurSOUKOS E. A. M. (1992) - Late Aptian to Maastrichtian foraminiferal biogeography andpalaeoceanogra-
phy of the Sergipe Basin, Brazil. Palaeogeogr., Palaeoclim., Palaeoecol., 92, p. 295-324.
KOursoUKOS E. A. M., DESTRO N., AzAMBUJA FILHO N. C. DE & SPADINI A. R. (in press) - Upper Aptian-Iower
Coniacian carbonate sequences in the Sergipe Basin, northeastern Brazil. In T. SIMO, B. W. SCOIT &
J.-P. MASSE (eds), Cretaceous Carbonate Platforms. Mem. Amer. Ass. Petrol. Geol.
KOurSOUKOS E. A. M. & HART M. B. (1990) - Cretaceous foraminiferal morphogroup distribution patterns,
palaeocommunities and trophic structures: a case study from the Sergipe Basin, Brazil. Trans. R. Soc.
Edinburgh, Earth Sci., 81, p. 221-246.
KOurSOUKOS E. A. M., MELLO M. R., AzAMBUJAFILHON. C. DE, HARTM. B. & MAxWELLJ. R. (1991a) - The
upper Aptian-Albian succession of the Sergipe Basin, Brazil : an integrated paleoenvironmental
assessment Amer. Ass. Petrol. Geol. Bull., 75, 3, p. 479-498.
KOurSOUKOS E. A. M., MELLO M. R. & AzAMBUJA FILHO N. C. DE (1991b) - Micropalaeontological and geo-
chemical evidence of mid-Cretaceous hypoxic-anoxic environments in the Sergipe Basin, northeas-
tern Brazil. I n R. V. TYSON & T. H. PEARSON (eds), Modern and ancient continental shelfanoxia. Geol.
Soc. Spec. Publ., 58, p. 427-447.
LANA M. C. (1990) - Bacia de Sergipe-Alagoas: uma hip6tese de tectono-sedirnentar.ln G. P. R.
GABAGLIA & E. J. M!LANr (eds), Origem e de bacias sedimentares, p. 311-332. Ed. Gavea:
R. Redisch Prog. Visual Prod. Graf. e : Petrobras, Rio de Janeiro.
MASTERS B. A. (1977) - Mesowic planktonic Foraminifera: a world-wide review and analysis. In RAMSAY A.
T. S. (ed.), Oceanic Micropal., I, p. 301-731. Academic Press, London.
MAURY C. J. (1937) - 0 Cretaceo de Sergipe. Serv. Geol. Miner. Brasil, Monogr., 11, p. 1-283.
OJEDA H. A. O. (1982) - Structural framework, stratigraphy and evolution of Brazilian marginal basins. Bull.
Amer. Ass. Petrol. Geol., 66, 6, p. 732-749.
OJEDA H.A. O. & FUGrrAA. M. (1976) - BaciaSergipe/Alagoas : Geologiaregional e perspectivas petroliferas.
Anais XXVIII Congr. (Brasil. Geol.) [porto Alegre, RS], I, p. 137-158.
PARRISH J. T. & CURTIS R. L. (1982) - Atmospheric circulation, upwelling, and organic-rich rocks in the
Mesozoic and Cenowic Eras. Palaeogeogr., Palaeoclim., Palaeoecol., 40,1-3: p. 31-66..
PETRI S. (1962) - Foraminiferos cretaceos de Sergipe. Bol. Fac. Filos., Cienc. Letras, Univ. Sa.o Paulo, 265
(Geologia, 20), p. 1-140.
PONTE F. C. & ASMUS H. E. (1976) - The Brazilian marginal basins: current state of knowledge. In ALMEIDA
F. F. M. DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc., 48, Suplem.,p. 215-
239.
261
RAsPLUS L. et al. (1987) - Stratigraphie integree du Sillon Citrabetique (Sierra de Fontealent, Province
d' Alicante, Espagne). Geobios, 20, 3, p. 337-387.
REYMENT R. A. & TAIT E. A. (1972) - Biostratigraphical datingof the early history ofthe South Atlantic Ocean.
Phil. Trans. R. Soc. London, B. Bioi. Sci., 264, 858, p. 55-95.
REYMENTR. A., BENGfSONP. & TAIT E. A. (1976) - Cretaceous transgressions in Nigeriaand Sergipe-Alagoas
(Brazil). In ALMEIDA F.P. M.DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc.,
48, Suplem., p. 253-264.
ROBASZ\'NSKI F. & CARON M. (eds) (1979) - Atlas de Foraminiteres planctoniques du Cretaee moyen (Mer
boreaIe et Tethys), Premiere partie. Cah. Micropal., 1979, 1, p. 1-185 ; Deuxieme partie. Cah.
Micropal., 1979,2, p. 1-181.
ROBASZ\'NSKIF., CARON M., GoNZALEs DoNOSOJ. M. & WONDERS A. A. H. (1984) - Atlas ofLateCretaeeous
Globotruncanids. Rev. Micropal., 26, 3-4, p. 1-305.
SCHAILER H. (ed.) (1970) -RevisAoestratigrMica da Baciade Sergipe/Alagoas. Bol. Tee. Petrobrds, 12, 1[for
1969], p. 21-86.
SCHAILER H., DEllA FAVERA J. C. & TIBANA P. (1980) - Roteiro Geol6gico da Bacia Sergipe-Alagoas.
Unpublished internal report, Petrobnis, Rio de Janeiro.
SMl1lI A. B. (1991) - Taxonomic descriptions. In A. B. SMl1lI & P. BENGfSON, Cretaceous echinoids from
north-eastern Brazil. Fossils and Strata, 31, p. 15-64.
SMl1lI A. B. & BENGfSON P. (1991) - Cretaceous echinoids from north-eastern Brazil. Fossils and Strata, 31,
p.1-88.
VAN HINTEJ. E. (1976) - ACretaceous time scale. Amer.Assoc. Petrol. Geol. Bull., 60,4, p. 498-516.
262
RAsPLUS L. et al. (1987) - Stratigraphie integree du Sillon Citrabetique (Sierra de Fontealent, Province
d' Alicante, Espagne). Geobios, 20, 3, p. 337-387.
REYMENT R. A. & TAITE. A. (1972) - Biostratigraphical datingof the early history ofthe South Atlantic Ocean.
Phil. Trans. R. Soc. London, B. Bioi. Sci., 264, 858, p. 55-95.
REYMENTR. A., BENGfsONP. & TAITE. A. (1976) - Cretaceous transgressions in Nigeriaand Sergipe-Alagoas
(Brazil). In ALMEIDA F.P. M.DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc.,
48, Suplem., p. 253-264.
ROBASZ\'NSKI F. & CARON M. (eds) (1979) - Atlas de Foraminiferes planctoniques du Cretace moyen (Mer
boreaIe et Tethys), Premiere partie. Cah. Micropal., 1979, 1, p. 1-185 ; Deuxieme partie. Cah.
Micropal., 1979,2, p. 1-181.
ROBASZ\'NSKIF., CARON M., GoNZALES DoNOSoJ. M. & WONDERS A. A. H. (1984) - Atlas of Late Cretaceous
Globotruncanids. Rev. Micropal., 26, 3-4, p. 1-305.
SCHAlLER H. (ed.) (1970) -RevisAoestratigrMica da Baciade Sergipe/Alagoas. Bol. Tee. Petrobrds, 12, 1[for
1969], p. 21-86.
SCHAlLER H., DEllA FAVERA J. C. & TIBANA P. (1980) - Roteiro Geol6gico da Bacia Sergipe-Alagoas.
Unpublished internal report, Petrobnis, Rio de Janeiro.
SMl1lI A. B. (1991) - Taxonomic descriptions. In A. B. SMl1lI & P. BENGfSON, Cretaceous echinoids from
north-eastern Brazil. Fossils and Strata, 31, p. 15-64.
SMl1lI A. B. & BENGfSON P. (1991) - Cretaceous echinoids from north-eastern Brazil. Fossils and Strata, 31,
p.1-88.
VAN HINTEJ. E. (1976) - ACretaceous time scale. Amer.Assoc. Petrol. Geol. Bull., 60,4, p. 498-516.
262
RAsPLUS L. et al. (1987) - Stratigraphie integree du Sillon Citrabetique (Sierra de Fontealent, Province
d' Alicante, Espagne). Geobios, 20, 3, p. 337-387.
REYMENT R. A. & TAIT E. A. (1972) - Biostratigraphical datingof the early history ofthe South Atlantic Ocean.
Phil. Trans. R. Soc. London, B. Bioi. Sci., 264, 858, p. 55-95.
REYMENTR. A., BENGfSONP. & TAIT E. A. (1976) - Cretaceous transgressions in Nigeriaand Sergipe-Alagoas
(Brazil). In ALMEIDA F.P. M.DE (ed.), Continental margins ofAtlantic type. AnaisAcad. Brasil. Cienc.,
48, Suplem., p. 253-264.
ROBASZ\'NSKI F. & CARON M. (eds) (1979) - Atlas de Foraminiteres planctoniques du Cretaee moyen (Mer
boreaIe et Tethys), Premiere partie. Cah. Micropal., 1979, 1, p. 1-185 ; Deuxieme partie. Cah.
Micropal., 1979,2, p. 1-181.
ROBASZ\'NSKIF., CARON M., GoNZALEs DoNOSOJ. M. & WONDERS A. A. H. (1984) - Atlas ofLateCretaeeous
Globotruncanids. Rev. Micropal., 26, 3-4, p. 1-305.
SCHAILER H. (ed.) (1970) -RevisAoestratigrMica da Baciade Sergipe/Alagoas. Bol. Tee. Petrobrds, 12, 1[for
1969], p. 21-86.
SCHAILER H., DEllA FAVERA J. C. & TIBANA P. (1980) - Roteiro Geol6gico da Bacia Sergipe-Alagoas.
Unpublished internal report, Petrobnis, Rio de Janeiro.
SMl1lI A. B. (1991) - Taxonomic descriptions. In A. B. SMl1lI & P. BENGfSON, Cretaceous echinoids from
north-eastern Brazil. Fossils and Strata, 31, p. 15-64.
SMl1lI A. B. & BENGfSON P. (1991) - Cretaceous echinoids from north-eastern Brazil. Fossils and Strata, 31,
p.1-88.
VAN HINTEJ. E. (1976) - ACretaceous time scale. Amer.Assoc. Petrol. Geol. Bull., 60,4, p. 498-516.
262