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morphology, and the riparian vegetation, and as a consequence, the structure of benthic
and fish communities (Ward and Stanford, 1983; Blinn and Cole, 1990; Stromberg and
Patten, 1990; Cazaunon and Giudicelli, 1999; Radar and Belish, 1999; Welcomme,
2001; Jacob et al., 2003; Uehlinger et al., 2003). Hydrodynamic responses to stream
regulation are primarily a function of alterations in the sediment load and the flow
regime and their interactions (Petts, 1980). Stream regulation, the anthropogenic
control of discharge of running waters, has profoundly influenced virtually all the
worlds major river systems (Ward and Stanford, 1979). Considerable literature on the
environmental modifications as a consequence of damming rivers is available (Neel,
1963; Srinivasan, 1963; Leopold et al., 1964; Ackerman et al., 1973; Rao and Palta,
1973; Dolan et al., 1974; Ridley and Steel, 1975; Ward, 1976, 1982; Baxter, 1977;
Buma and Day, 1977; Ward and Stanford, 1979; Mittal, 1981; Raju et al., 1981;
Leentvaar, 1985; Singit et al., 1988; Sehgal, 1978; Simon, 1979; Petts, 1980; Sharma,
1991; Moog, 1993; Camargo and Voelz, 1998; Cazaubon and Giudicelli, 1999; Radar
and Belish, 1999; Thomaz et al., 1999; Pringle et al., 2000; Sugunan, 2000; Wright et
al., 2000; Rolauffs et al., 2001; Vinson, 2001; Welcomme, 2001; Klemm et al., 2002;
Mishra et al., 2002; Jakob et al., 2003; Uehlinger et al., 2003; Pollard and Reed, 2004;
Chakrabarty and Das, 2005; Thomson et al., 2005; Bellingor et al., 2006 and Fumiko
et al., 2006). A considerable work has been done previously on the impact of Tehri
Dam construction on lotic environment of Bhagirathi during 1979-1987 (Sharma, 1981
and 1991). But since then no work has been done for the study of further changes
going on in the aquatic environment of fish habitats.
It is quite evident that the fish productivity of any aquatic ecosystem is
influenced by its physico-chemical and biological characteristics. The biota of high
altitude lotic environment is very sensitive to any change in the characteristics of
water. Therefore, for determining any depression in the biotic potential of high
altitude fluvial system, it is a prime requisite for having knowledge on the
environmental stress manifested in the form of an adverse change in the limnological
parameters imposed by human interference. Therefore, it was felt necessary to
undertake an assessment of ecological impact on aquatic environment of fish habitats
of Bhagirathi River.
OBSERVATIONS
The immediate consequence of the construction of a dam, whatever its size in
an aquatic ecosystem, is the modification of the hydrological regime, creating thus a
lentic environment and change the water flow and water quality downstream the dam.
Thus, the artificial lake upstream and drastically altered riverine environment
downstream, both these are unfavourable for the aquatic organisms including fish
communities inhabiting the natural ecosystem.
An assessment of transformation in riverine morphology and degradation of
fish habitats of Bhagirathi River caused due to Tehri Dam construction activities was
made for the entire period of construction. However, a regular monitoring was made
for two annual cycles (September 2004 - August 2006).
A. Geomorphic Transformations in Bhagirathi River
Variety in habitat is a consequence of the physical form of the river. Channels
show curvature, vary in their cross-sectional profile, alternate between riffles, pools
and other channel categories, exhibit much horizontal and vertical variation in current
velocity and contain a variety of bottom substrates, which often vary from place to
place in accord with channel and velocity. Its no wonder the fisheries biologists assess
habitat variability as a measure of stream health (Cushing and Allan, 2001).
One of most remarkable features of running water is its tendency to curve and
tracing a sinuous path. Riffles, regions of shallow, faster water; and pools, regions of
slower, deeper water, are characteristics of virtually all rivers. Substrate and channel
curvature play a role in the alteration of riffles and pools. In relatively straight, gravelbed river, a riffle-pool sequence tends to repeat at intervals of roughly 5-7 channel
widths. In very steep, boulder filled channels, steep pools are separated by riffles,
cascades, or small water falls. In streams containing substrate in variety of sizes, finer
substrate usually is found in the pools because slower currents there favour deposition
of suspended materials. Coarse substrate usually characterizes riffles, where steeper
gradient and faster current erode and transport finer particles, leaving behind the
larger cobbles and small boulders (Cushing and Allan, 2001).
Natural morphology of pools and riffles forms was altered drastically at many
places at the impacted site (S2). Due to the alteration in the substrate composition and
river curvature, the width of the river reduced and the flow increased at many points
at S2. The sequence of pool-riffle was changed. The riffles, rapids and cascades were
converted into dammed pools at some places. However, the wide white water pools
were converted into turbid and narrow pools at other places. These dam construction
activities have also resulted in the inundation of spawning and feeding grounds of
fish, choking of breeding grounds, blocking of migration channels of coldwater fish
and destruction of favourable habitats for aquatic benthic insects. Thus, a distinct
change in the composition of substrate was observed in the ecotransformed site (S2).
Few changes have also been observed in the course of river upstream between Dobata
(2 km upstream the dam) and the Tehri Dam. The river channel has also been changed
morphometrically from its natural course. The river has widened due to the digging
and leveling of the ground in close vicinity of the dam. The natural bottom substrata
have been replaced mainly by silt and small pebbles as the big boulders and cobbles
were extracted from this area for crushing into fine sand. Pools became dominant here
than the riffles. However, no change was observed at the reference site (S1) as it was
free from the impact of Tehri Dam construction activities. A major change in the river
morphology was resulted due to the closure of tunnels on October 29, 2005. The
entire area of fluvial system of Bhagirathi was converted into a huge lentic water body
(Fig. 5.2). A drastic change was also resulted downstream to Tehri Dam. The river
between Tehri and Deoprayag was reduced to small stream for two months after the
closure of tunnels (Fig. 5.3).
B. Degradation of Aquatic Environment
The physico-chemical profile of the lotic environment of Bhagirathi at the
reference site (S1) and impacted site (S2) has been portrayed over two-year period
(September 2004 - August 2006) in Table 5.1.The seasonal variations in the physicochemical parameters of aquatic environment at both the sampling sites (S1 and S2) has
been presented in the Tables 5.2 - 5.3.
a. Change in Physico-environmental Variables:
The atmospheric temperature recorded at both the sampling sites was almost
similar (S1: 21.8 5.2 0C and S2: 21.9 5.2 0C) with the maximum value (28.0 0.7
0
C) during monsoon season at S2 and minimum value (15.9 1.4 0C) during winter
season at S1. Water temperature recorded at both the sites with the higher value (15.6
2.8 0C) at S2 than S1 (15.0 3.1 0C). The water temperature was maximum (17.8
2.1 0C) during summer at S2 and minimum (11.3 0.5 0C) during winter at S1. A
slight change in the water temperature at S2 was noticed. It may be due to mixing of
muck in the water and removal of riparian vegetation. The water temperature of
Bhagirathi gradually increases from January to April and thereafter, shows a declining
trend upto the month of August, because of the melting of snow at the higher
elevations from where the river originates (Sharma, 1984 e).
Water current of Bhagirathi is turbulent and mainly of shooting type. A slight
difference was noticed in the water current caused by dam construction activities. It
was recorded higher (2.37 1.31 m sec-1) at S2 than S1 (2.04 0.88 m sec-1). The
maximum value (4.20 0.21 m sec-1) of water current was recorded in monsoon
months (July-August) and minimum (0.97 0.41 m sec-1) in winter months
(NovemberFebruary) at S2.
A minor change in the hydromedian depth was also recorded. The maximum
value (2.37 0.38 m) of HMD was observed during monsoon and minimum value
(0.55 0.19 m) during winter.
A considerable difference was observed in the turbidity at S2 due to dam
construction. It was recorded 160.61 239.09 NTU at S2 and 138.13 244.75 NTU at
S1. Seasonal data of turbidity revealed that the maximum turbidity (663.25 215.31
NTU) was recorded in monsoon and minimum (7.13 4.15 NTU) in winter.
Transparency was also influenced by the dam construction activities. It reduced from
0.35 0.29 m (S1) to 0.21 0.17 m (S2). Seasonally, the transparency was maximum
(0.62 0.15 m) during winter and minimum (0.01 0.01 m) during monsoon.
Conductivity is a measure of the total amount of ions present in a body of
water and is a useful approximation to chemical richness, was also influenced by the
anthropogenic perturbations (dam construction activities) at S2 (0.16 0.06 S cm-1).
It was (0.13 0.01 S cm-1) in a natural environment at S1. Seasonal data of
conductivity revealed that the maximum conductivity (0.16 0.05 S cm-1) was
recorded during summer (March-June) and minimum (0.05 0.03 S cm-1) during
winter (November-February).
A considerable change in total dissolved solids (TDS) was observed at S2
(45.14 30.13 mg l-1) than S1 (34.0 33.39 mg l-1). The maximum TDS (100.0
18.38 mg l-1) was observed during the monsoon season and minimum (8.25 3.71
mg l-1) during winter.
b. Change in Chemico-environmental Variables:
A significant change in the dissolved oxygen as a consequence of dam
construction activities was recorded at S2 (9.79 0.50 mg l-1) than the reference site
(10.55 0.84 mg l-1). The dissolved oxygen was recorded highest (11.17 0.72
mg l-1) during winter and lowest (9.23 0.13 mg l-1) during monsoon. In Bhagirathi
River, the higher concentration of dissolved oxygen was observed in the colder
months (November-February), followed by a sharp decline in the monsoon months
(July-August). Free Carbon dioxide showed the inverse relationship with the
dissolved oxygen. Free CO2 was maximum (3.52 0.16 mg l-1) during wet months
and minimum (1.63 0.40 mg l-1) during the colder months.
A minor change in the annual means of hydrogen ion concentration (pH) was
recorded at the impacted site (7.63 0.17) than the reference site (7.51 0.18).
Hardness and alkalinity also followed the pH values. The higher value of hardness
was recorded at S2 (64.92 17.28 mg l-1) than S1 (59.88 16.16 mg l-1). Alkalinity
was recorded with the higher value (53.67 6.03 mg l-1) at S2 than S1 (49.67 6.08
mg l-1).
Chlorides were recorded higher at S2 (3.18 0.70 mg l-1) than S1 (2.61 0.56
mg l-1) with the maximum value (4.24 0.05 mg l-1) during monsoon and minimum
(2.24 0.25 mg l-1) during winter.
The concentrations of nitrates and phosphates were also influenced by the dam
construction activities. These concentrations slightly increased at the impacted site
(0.48 0.25 mg l-1 ; 0.43 0.45 mg l-1) than the reference site (0.41 0.24 mg l-1 ; 0.25
0.34 mg l-1).
c. Depression in Biological Variables:
Impact on the biological variables of the fluvial system of Bhagirathi at the
impacted site (S2) in relation to reference site (S1) has been presented in the Table 5.4.
The biotic components of fish habitat of the Bhagirathi River are contributed
by periphyton-phytoplankton, zooplankton and macroinvertebrates. The density of
periphyton-phytoplankton, zooplankton and macroinvertebrates was recorded 530.54
285.80 units l-1 ; 50.50 7.78 units l-1 and 895.83 623.70 units m-2 respectively at
the reference site (S1). The density of these biological components - periphytonphytoplankton, zooplankton and macroinvertebrates were reduced drastically to 255.0
126.38 units l-1 ; 35.0 4.24 units l-1 and 172.92 171.45 units m-2 respectively at
the impacted site (S2) as a consequence of several civil construction activities.
The periphyton generally found attached to rocks, submerged wood and
floating or submerged vegetation and were abundant in the fast current of shallow
riffles. The phytoplankton were found in the slower current of deeper pools. Both
these components contribute the primary trophic level and were mainly represented
by
Bacillariophyceae,
Chlorophyceae
and
Myxophyceae.
The
periphyton-
monsoon
(Table
5.10).
Beta
diversity
was
also
calculated
for
variables,
substrate
composition
and
geomorphometric
works and other anthropogenic perturbations (Mills et al., 1966; Canton and Ward,
1978; Nilsson, 1978; Cline et al., 1982, Leentvaar, 1985 and Sharma, 1991).
Cline et al., (1982) reported the distinct deleterious effects of highway
construction on the macroinvertebrates and epilithic algae of a high mountain stream
in Northern Colorado. According to them suspended solids and the proportion of fine
sediment in the substrate increased at impacted sites. At impacted sites algal species
diversity and the organic content of the epilithon were reduced. The macroinvertebrate
community was altered by construction activities. Reduction in density, abundance and
diversity were apparent and the taxonomic composition was modified. Leentvaar
(1985) reported an account of chemistry and biological data of the Sinu river system in
Western Columbia and the consequence of inundation of tropical rain forest by Alto
Sinu hydroelectric project. A sharp reduction in the density of diatoms, blue-green
algae and nymphs of aquatic macroinvertebrates was noticed. Mills et al., (1966) made
an attempt on the distinct changes in the physico-chemical environment and the biota
inhabiting it as a consequence of anthropogenic perturbations. The effects of the
construction of a motorway in Essex, U.K. on the lotic environment of a stream
receiving impact were studied by Extence (1978). He noticed a drastic change in the
suspended solids and substrate composition and a deleterious effect of the modified
environment on stoneflies, mayflies and cased caddisflies inhabiting the stream.
The temperature being an important factor in aquatic environments regulates
the composition of biotic profile as well as other characteristics of water. It also exerts
profound direct or indirect influence on metabolic and physiological behaviour of
aquatic ecosystem (Welch, 1952), which reflects in the dynamics of living organisms.
Alterations in the water velocity, temperature and dissolved oxygen are
usually found downstream from the main dam site (Ward and Stanford, 1979; Petts,
1984; Craig and Kemper, 1987; Gore and Kemper, 1987; Moog, 1993; Camargo and
Voelz, 1998 and Camargo et al., 2004). A significant alteration in the water velocity,
temperature and dissolved oxygen was also found downstream at the impacted site
(S2) of Tehri Dam under the present study. The same results were also observed by
Rolauffs et al. (2001) while studying the invertebrate composition and productivity of
Beaver Dam in Germany. The concentration of phosphates and nitrates was also
observed high at the impacted site than the reference site. According to Fumiko et al.,
1994; Wright et al., 2000 and Klemm et al., 2002). Chutter (1969) reported a
deleterious effect of sand and silt deposition on macroinvertebrates of South African
streams. Substantive alterations in the substrate composition and transportation of fine
sediments filling the interstitial spaces have been noticed at the impacted site (S2) due
to Tehri Dam construction under the present study.
Nilsson (1978) made an attempt to use floristic analysis to monitor the
vegetational changes along a stretch of the Swedish river, Umealven, following setting
up of hydroelectric works. Changes in silting, water flow and water level amplitude
were noted. The changes have brought alterations in the abundance and distribution of
some species, few have disappeared or immigrated.
Impacted Site (S 2)
Environmental Variables
X
S.D.
S.D.
21.8
5.2
21.9
5.2
15.0
3.1
15.6
2.8
2.04
0.88
2.37
1.31
HMD (m)
1.21
0.72
1.16
0.70
138.13
244.75
160.61
239.09
0.35
0.29
0.21
0.17
Conductivity ( S cm )
0.13
0.01
0.16
0.06
34.0
33.39
45.14
30.13
10.55
0.84
9.79
0.50
1.90
0.42
2.79
0.60
pH
7.51
0.18
7.63
0.17
59.88
16.16
64.92
17.28
49.67
6.08
53.67
6.03
2.61
0.56
3.18
0.70
0.41
0.24
0.48
0.25
0.25
0.34
0.43
0.45
Air temperature ( C)
0
Water temperature ( C)
-1
Turbidity (NTU)
Transparency (m)
-1
-1
Chlorides (mg l )
-1
Nitrates (mg 1 )
-1
Phosphates (mg l )
Autumn
(Sep-Oct)
23.7 3.5
Winter
(Nov-Feb)
15.9 1.4
Summer
(March-Jun)
23.7 3.8
Monsoon
(Jul-Aug)
27.9 0.5
15.4 1.2
11.3 0.5
17.5 2.0
16.9 0.3
2.44 0.29
1.64 0.72
1.66 0.89
3.22 0.10
HMD (m)
1.53 0.13
0.65 0.19
1.02 0.52
2.37 0.38
Turbidity (NTU)
12.25 8.13
7.13 4.15
84.50 82.24
633.25 189.15
0.40 0.39
0.62 0.15
0.20 0.11
0.04 0.03
0.14 0.03
0.13 0.05
0.15 0.04
0.05 0.03
20.50 2.12
8.25 3.71
33.25 21.75
87.00 2.83
10.24 0.36
11.17 0.72
10.54 0.79
9.66 0.23
2.09 0.16
1.63 0.40
1.79 0.25
2.50 0.01
pH
7.50 0.04
7.33 0.10
7.56 0.10
7.79 0.06
63.25 20.15
72.0 7.0
57.25 4.62
37.50 3.54
45.75 8.84
52.50 2.86
51.63 4.48
44.00 5.66
2.88 0.28
2.24 0.25
2.38 0.12
3.54 0.22
0.46 0.04
0.18 0.14
0.41 0.16
0.79 0.04
0.09 0.12
0.10 0.09
0.33 0.49
0.56 0.30
-1
Transparency (m)
-1
Conductivity ( S cm )
-1
TDS (mg l )
-1
-1
Hardness (mg l )
-1
Alkalinity (mg l )
-1
Chlorides (mg l )
-1
Nitrates (mg 1 )
-1
Phosphates (mg l )
Autumn
(Sep-Oct)
23.9 3.3
Winter
(Nov-Feb)
16.2 1.2
Summer
(March-Jun)
23.7 3.8
Monsoon
(Jul-Aug)
28.0 0.7
15.5 1.1
12.1 1.1
17.8 2.1
17.1 0.2
2.56 0.44
0.97 0.41
2.53 1.05
4.20 0.21
HMD (m)
1.62 0.26
0.55 0.19
0.97 0.42
2.19 0.28
53.25 23.69
41.21 32.52
96.38 89.15
663.25 215.31
0.13 0.01
0.14 0.08
0.18 0.12
0.01 0.01
0.14 0.04
0.15 0.03
0.16 0.05
0.07 0.02
37.50 3.54
25.42 2.79
38.50 20.29
100.0 18.38
9.50 0.32
10.18 0.52
9.90 0.36
9.23 0.13
2.97 0.31
2.35 0.41
2.73 0.57
3.52 0.16
pH
7.73 0.11
7.45 0.08
7.62 0.08
7.84 0.01
71.00 16.97
75.25 12.42
61.25 7.64
45.75 1.77
53.75 5.30
55.17 4.18
56.25 5.66
45.50 4.24
3.24 0.30
2.66 0.49
3.08 0.40
4.24 0.05
0.40 0.32
0.27 0.09
0.51 0.08
0.85 0.08
0.20 0.08
0.18 0.09
0.54 0.61
0.84 0.11
-1
Turbidity (NTU)
Transparency (m)
-1
Conductivity ( S cm )
-1
TDS (mg l )
-1
-1
Hardness (mg l )
-1
Alkalinity (mg l )
-1
Chlorides (mg l )
-1
Nitrates (mg 1 )
-1
Phosphates (mg l )
Table 5.4. Comparative biological profi le of fish habitats of Bhagirathi at reference site (S 1 )
and impacted site (S 2)
Environmental
Variables
Reference Site (S 1)
X
Impacted Site (S 2 )
S.D.
S.D.
183.88
66.85
35.07
285.80
195.50
41.0
18.50
255.0
88.17
25.56
12.64
126.38
3.54
1.41
0.0
2.83
7.78
4.50
18.50
6.50
5.50
35.0
2.12
0.71
0.71
0.71
4.24
331.85
96.49
85.64
31.79
28.78
17.37
15.57
16.28
623.70
96.67
23.13
41.25
2.50
2.92
2.50
1.04
2.92
172.92
71.82
28.79
44.49
9.73
1.86
3.61
3.61
7.53
171.45
Table 5.5. Seasonal variations in density of biological components (fish food) of Bhagirathi
at reference site (S 1)
Environmental
Variables
Autumn
(Sep-Oct)
Winter
(Nov-Feb)
Summer
(March-Jun)
Monsoon
(Jul-Aug)
278.50 38.89
517.13 131.64
391.50 146.76
105.0 55.15
Chlorophyceae
49.50 0.71
154.75 64.75
136.75 41.83
33.0 32.53
Myxophyceae
19.0 9.90
71.50 24.22
70.50 30.38
14.0 9.90
347.0 49.50
743.38 220.61
598.75 214.97
152.0 97.58
Zooplankton (units l )
Cladocera
2.0 0.71
7.0 0.70
2.50 0.77
0.0 0.0
Protozoa
2.0 0.71
11.0 1.22
6.00 0.50
2.0 0.0
Copepoda
0.50 0.35
4.50 1.46
2.50 0.54
0.50 0.35
0.0 0.0
6.50 0.77
3.50 0.54
0.0 0.0
4.50 1.77
29.0 4.14
14.50 2.34
2.50 0.35
Total
-1
Rotifera
Total
-2
Macroinvertebrates (units m )
Ephemeroptera
50.0 16.70
95.70 44.98
45.31 20.12
6.94 0.98
Trichoptera
31.25 8.84
88.19 34.97
36.46 13.36
18.75 5.89
Diptera
32.81 2.21
76.82 31.31
39.06 16.63
20.31 4.42
Coleoptera
30.0 3.54
18.75 9.95
22.50 17.44
12.50 1.77
Hemiptera
37.50 0.0
26.56 18.41
20.31 25.65
0.0 0.0
Plecoptera
12.50 0.0
17.19 11.84
7.81 0.0
3.13 0.0
Odonata
25.0 0.0
8.33 0.0
12.50 0.0
0.0 0.0
Mollusca
0.0 0.0
15.63 6.25
12.50 11.97
0.0 0.0
219.06 31.28
347.18 157.71
196.46 105.16
61.63 13.06
Total
Table 5.6. Seasonal variations in the density of biological components (fish food) of
Bhagirathi at the impacted site (S2)
Environmental
Winter
(Nov-Feb)
Summer
(March-Jun)
Monsoon
(Jul-Aug)
131.50 26.16
268.25 70.09
213.0 62.52
79.0 22.63
Chlorophyceae
21.0 9.90
65.0 18.27
43.0 16.73
9.0 2.83
Myxophyceae
7.0 4.24
24.0 9.94
25.0 11.09
6.0 2.83
357.25 98.30
281.0 90.35
94.0 28.28
Variables
Autumn
(Sep-Oct)
Total
159.50 40.31
Zooplankton (units l-1)
Cladocera
0.0 0.0
3.0 0.29
1.50 0.54
0.0 0.0
Protozoa
1.0 0.71
10.0 1.11
7.0 0.91
0.50 0.35
Copepod
0.50 0.35
3.0 0.73
3.0 0.73
0.0 0.0
Rotifera
0.0 0.0
4.0 0.50
1.50 0.54
0.0 0.0
1.50 1.06
20.0 2.63
13.0 2.71
0.50 0.35
Total
12.22 3.93
18.01 9.43
8.47 5.14
1.25 0.0
Trichoptera
14.17 2.95
28.13 5.49
6.67 7.61
0.0 0.0
Diptera
18.75 11.05
10.31 5.19
11.15 6.56
3.44 0.0
Coleoptera
0.0 0.0
0.0 0.0
1.25 0.0
0.0 0.0
Hemiptera
0.0 0.0
0.0 0.0
0.0 0.0
0.0 0.0
Plecoptera
3.13 0.0
1.25 1.44
0.0 0.0
0.0 0.0
Odonata
0.0 0.0
0.0 2.13
0.0 0.0
0.0 0.0
Mollusca
0.0 0.0
4.38 5.91
0.0 0.0
0.0 0.0
48.26 17.92
62.08 31.08
27.53 19.32
4.69 0.0
Total
Table 5.7. Annual mean of density of periphyton -phytoplankton recorded at reference site (S 1) and impacted site (S 2)
over a period of two-year (September 2004 -August 2006)
Year
Site
Bacillariophyceae
Chlorophyceae
Myxophyceae
Total
S1
380.08 174.56
110.83 71.61
53.5 34.58
544.42 281.01
S2
353.50 193.20
111.0 62.01
52.17 35.29
516.67 290.58
S1
199.17 84.58
41.50 27.65
17.0 13.97
257.67 126.21
S2
191.83 91.76
40.50 23.48
20.0 11.31
252.33 126.56
2004-05
2005-06
Table 5.8. Annual mean of density of zooplankton recorded at reference site (S 1) and impacted site (S 2 ) over a period of
two-year (September 2004 -August 2006)
Year
Site
Cladocera
Protozoa
Copepoda
Rotifera
Total
S1
14.0 1.03
22.0 1.11
8.0 0.78
8.0 0.78
52.0 3.81
S2
6.0 0.52
18.0 1.31
7.0 0.67
6.0 0.67
37.0 3.18
S1
9.0 0.75
20.0 1.07
8.0 0.98
12.0 0.95
49.0 3.77
S2
3.0 0.45
19.0 1.16
6.0 0.67
5.0 0.51
33.0 2.81
2004-05
2005-06
Table 5.9. Annual mean of density of macroinvertebrates recorded at reference site (S 1) and impacted site (S2 ) over a period of two-year
(September 2004-August 2006)
Year
Site
Ephemeroptera
Trichopte
ra
Diptera
Coleoptera
Hemiptera
Plecoptera
Odonata
Mollusca
Total
S1
479.17
378.37
135.42
77.36
154.17
100.47
60.42
34.47
37.50
27.18
25.0
18.46
4.17
14.43
6.25
15.54
902.08
666.29
S2
110.42
72.66
16.67
18.63
33.33
34.27
0.0
0.0
5.0
9.77
3.33
8.07
2.08
7.22
2.92
7.53
173.75
158.15
S1
452.08
285.33
158.33
115.62
143.75
70.81
64.58
29.11
31.25
30.39
16.67
16.28
10.42
16.71
12.50
16.85
889.58
581.11
S2
82.92
60.73
29.58
28.00
49.17
39.82
5.0
14.46
0.83
2.89
1.67
3.89
0.0
0.0
2.92
7.53
172.09
157.31
2004-05
2005-06
Table 5.10. Seasonal variations in alpha diversity of primary and secondary trophic levels of
Bhagirathi River at both the sampling sites
Autumn
(Sep-Oct)
Winter
(Nov-Feb)
Summer
(March-Jun)
Monsoon
(Jul-Aug)
S1
40
40
40
32
S2
16
22
25
13
28
31
32.5
22.5
S1
S2
4.5
8.5
7.5
2.5
S1
22
27
27
12
S2
10
13
10
16
20
18.5
7.5
2004-06
Periphyton-phytoplankton
Zooplankton
Macroinvertebrates
Table 5.11. Beta diversity at the primary and secondary trophic levels of Bhagirathi River at both
the sampling sites
Year 2004-06
Average no. of
species
Beta diversity
32.5
1.231
8.5
1.058
20.0
1.350
Periphytonphytoplankton
S1
S2
40
25
Zooplankton
S1
S2
Macroinvertebrates
S1
27
S2
13
Autumn
(Sep-Oct)
S1
S2
Winter
(Nov-Feb)
S1
S2
Summer
(March-Jun)
S1
S2
Monsoon
(Jul-Aug)
S1
S2
++
++
+
+
++
+
+
+
+
++
+
+
++
+
+
+
+
+
+
+
++
+
+
+
+
++
+
++
+
++
+
++
+
+
+
+
+
+++
+++
++
+
+++
+
+
++
+
+++
++
+
+++
++
++
++
++
+
++
++
++
+
++
++
++
+++
++
+
+++
+
+++
++
+++
+
+
+
+
+
+
+
+
++
+++
+
+
+++
+
+
++
+
+++
+
+
+++
+
+
+
+
+
+
++
+
+
+
+
+
++
++
+
++
+
++
+
++
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
+
-
+
++
+
+
+
+
+
+
+
+
+
+
+
+
-
++
++
++
+
+
++
++
++
++
++
++
+
+
+
+
+
-
+
+
++
+
+
++
++
+
+
++
+
+
+
+
+
+
-
+
+
+
+
+
+
+
+
+
+
-
+
+
+
+
++
++
++
+
++
+
+
+
+
+
+
++
++
34
B. Zooplankton
B.1 Cladocera
Daphnia
Ceriodaphnia
B.2 Protozoa
Vorticella
Zoothamnium
B.3 Copepoda
Cyclops
Diaptomus
B.4 Rotifera
Trichocera
Asplanchna
Keratella
C. Macroinvertebrates
C.1 Ephemeroptera
Heptagenia
Leptophlebia
Caenis
Cloeon
Baetis
Centroptilum
Ephemerella
Ecdynurus
Siphlonuridae
C.2 Trichoptera
Hydropsyche
Rhyacophila
Glossosoma
++
++
++
+
++
+
+
+++
++
++
++
++
+
++
+
+
+
+
-
++
++
++
++
++
+
+
+
+
+
++
++
+++
++
+++
++
+
++
+
++
+
+
+
-
+
+
++
+
-
+++
+++
++
+++
++
++
+
++
++
+
++
+
++
+
-
++
++
+
++
+
++
+
+
+
++
+
-
+
+
+
-
+
-
++
++
+++
++
+++
++
++
++
++
+++
++
+++
++
+
-
+
-
++
+
+
+
-
++
+
+
+
+
-
+
-
+
+
+
++
+
++
++
+
+
+
++
++
++
+
+
+
-
+
+
+
-
++
++
C.3 Diptera
Tendipes
Tabanus
Antocha sp.
Psychoda
C.4 Coleoptera
Ochthebius
Dineutus sp.
Amphizoa lecontei
Agabinus glabrellus
Peltodytes
C.5 Hemiptera
Corexia
Hesperocorixa
C.6 Plecoptera
Brachyptera
Acroneuria
C.7 Odonata
35
Hagenius
+
C.8 Mollusca
Lymnaea
+++ Abundant; ++ Common; + Rare; - Absent
++
++
36
500
S1
S2
-1
Density (ind l )
400
300
200
100
0
eae
hyc
l
c il
Ba
op
ari
eae
hyc
p
o
lor
Ch
ph
yxo
ae
yce
S1
160
S2
-1
Density (ind l )
140
120
100
80
60
40
20
0
Cladocera
Protozoa
Copepoda
Rotifera
S1
S2
-2
Density (ind m )
500
400
300
200
100
0
a
a
a
ta
ra
ra
ra
ca
te r
te r
ter
pte
pte c opte dona ollus
rop i chop
Dip ole o
mi
O
e
e
e
l
M
H
P
C
Tr
he m
Ep
37
Fig. 5.1. Fundamental pool and riffle forms prevalent in mountain fluvial system of Garhwal
Himalayas
38
Fig. 5.2. Morphometric transformation of Bhagirathi fluvial system into a huge lenti c
water body after impoundment