Professional Documents
Culture Documents
Habitat Selection by Stream Indicator Biota: Development of Biological Tools for the
Implementation of Protective Minimum Flows for Florida Stream Ecosystems
i
ACKNOWLEDGEMENTS
The authors extend their gratitude to James Gore, Ph.D., of the University of South
Florida at St. Petersburg, for his invaluable contributions to this project, which included selection
curves, execution of the PHABSIM model for a number of taxa and community quality
The authors also express their gratitude to John Good, Ph.D., of the Suwannee River
Water Management District, who graciously provided an historical flow simulation for the
We thank the following individuals from the University of Florida who assisted us with
field efforts: Jason Bennett, Doug Colle, Drew Dutterer, and Lauren Marcinkiewicz.
We thank the following present and former FWC staff for their dedicated efforts in the
field and/or laboratory: Tracey Smith, John Johansen, Cara Miller, Ann Cichra, and Amanda
Waddle.
The authors thank the U.S. Fish and Wildlife Service for providing funding for this work
through the state wildlife grant program, and Brian Branciforte of the FWC Habitat and Species
Conservation Division for his administrative efforts associated with the project.
We also thank Dick Krause and Jim Estes of FWC for their reviews of the initial draft.
ii
TABLE OF CONTENTS
Abstract ..............................................................................................................................................i
Acknowledgements ......................................................................................................................... ii
Introduction ......................................................................................................................................1
Background ......................................................................................................................................2
Objectives.........................................................................................................................................6
Study Area........................................................................................................................................7
Results ............................................................................................................................................20
Discussion ....................................................................................................................................106
Conclusions ..................................................................................................................................112
Appendices...................................................................................................................................120
iii
Habitat Selection by Stream Indicator Biota: Development of Biological Tools for the
Implementation of Protective Minimum Flows for Florida Stream Ecosystems
INTRODUCTION
stewardship in the midst of development and increasing water use by a growing human
population. From 1950 to 2005 Floridas population grew from 2.8 to 17.9 million, a 539 %
increase (Smith 2005). During this same period, withdrawals of freshwater from surface and
groundwater sources increased from 1,454 million gallons per day to 6,868 million gallons per
day (372 % increase; Marella 2008). Withdrawals of groundwater have exceeded aquifer
feet (Berndt et al 1998) and reduced surface water flows (Vergara 1994, Rao and Clapp 1996,
Berndt et al. 1998). Groundwater pumping has been responsible for complete cessation of spring
flows (e.g. Kissengen Spring, Polk Co.; Berndt et al. 1998). As Floridas population continues to
grow and land development progresses, freshwater withdrawals from surface and underground
sources can be expected to increase (Vergara 1994, Rao and Clapp 1996, Berndt et al. 1998), and
competition for the dwindling resource among household users, industry, and agriculture will
Streams are complex and dynamic systems that are affected by a multitude of physical
and biological variables. Flow and water level are among the most ecologically influential of
these variables (Hynes 1970, Allan 1995). The magnitude, frequency, duration, timing, and rate
of change of flow events that characterize the natural flow regime influence the water quality,
energy resources, physical habitat and biotic interactions that collectively determine the
ecological integrity of lotic systems (Resh et al. 1988, Poff and Ward 1989, Poff et al. 1997,
1
Richter et al. 1997). Alteration of the natural flow regime, such as reduction of flows to levels
chronically below the normal hydrological range, pose a substantial threat to the integrity of
stream ecosystems (Poff and Ward 1989, Richter et al. 1997). Abiotic factors such as dissolved
oxygen, temperature, suspended sediments, bed load, and streambed stability are directly
influenced by flow and, in turn, directly affect the distribution of stream-dwelling organisms
(Ward and Stanford 1983, Allan 1995, Richter et al. 1996). Many stream organisms (e.g. net-
spinning caddisflies) are directly dependent upon specific ranges of flow for survival (Hynes
1970, Statzner et al. 1988, Gore et al. 2001). Physical habitat alterations caused by
anthropogenic stream flow manipulation have been shown to influence the abundance and
species composition of fishes in cold and warm water streams (Schlosser 1985; Bain et al. 1988;
Given the likely future increases in groundwater and surface water withdrawals, and the
potential for ecological damage resulting from these withdrawals, natural resource agencies and
prepared to provide constructive input to the water management districts at the outset of the MFL
establishment process. Insights provided by these additional inputs will help ensure the
formulation of flow recommendations that attain the overall goal of sustaining the ecological
integrity of streams. The present project was implemented as an initial step in this process.
Background
Floridas five water management districts are required by law (Section 373.042 FS) to
establish MFLs for streams and lakes within their respective jurisdictions. The statute defines
minimum flow as the limit at which further withdrawals would be significantly harmful to the
water resources or ecology of the area. The law further states that minimum flows and levels
2
are to be determined using the best available information and are to reflect seasonal variation
when appropriate. The statute provides no specific guidance regarding the terms significantly
harmful and best information available, but State Water Policy (62-40.473 F.A.C.) requires
that the water management districts must consider all of the following factors when determining
MFLs:
The overall intent of the legislation and policy directive was to provide adequate water
for consumptive users while ensuring maintenance of flows and levels necessary to protect biota
and safeguard ecological processes within streams, springs, and lakes. However, the concept of
regulating stream withdrawals using a minimum flows approach, which emerged in the
western U.S. states during the 1960s, has been substantially criticized for its inability to provide
for sustainable ecosystem function or habitat protection (Annear et al. 2004, Calow and Petts
1992, 1994). The Florida Fish and Wildlife Conservation Commission (FWC), as the state
agency charged with protecting Floridas fish, wildlife, and habitat resources, undertook the
work presented herein as an initial step in the formulation of biological tools intended to protect
stream-dwelling biota in the MFL process. FWC contributions are intended to support the water
management districts efforts to develop MFLs that are truly protective of aquatic ecosystems
3
Aquatic Inve rtebrate and Fish Communities as Ecological Indicators
secondary producers invertebrates act as a fulcrum in aquatic food webs by converting the
products of primary production into carbohydrate and protein for export up the food chain.
Invertebrates are a primary food resource to many fish, amphibian, reptile, bird, and mammal
species, as well as to other invertebrates. Invertebrates also play critical roles in the
decomposition and nutrient cycling processes via their feeding and digestive activities. Adverse
health. Invertebrate community structure has been used as a tool to evaluate the status of
freshwater systems since the origin of the science of limnology in the early 20th century
(Thienemann 1925, Brundin 1949, Brinkurst 1974). Early researchers recognized that
assemblages are essentially sessile and unable to escape disturbance, they are sculpted by, and
reflect, the habitat conditions in which they develop. Consequently, invertebrate community
structure, and the presence/absence and abundance of certain key invertebrate taxa, function as
structuring invertebrate communities. Bovee et al. (1978) demonstrated that invertebrates have
narrower tolerances of flow change than fish. Many stream-dwelling invertebrate taxa require
flow (as current velocity) for survival. Flowing water supplies food to gathering and filter-
feeding taxa and oxygen to taxa that respire via gills and spiracles. Among those taxa most
dependent upon flows are many species of mayflies (Ephemeroptera), stoneflies (Plecoptera),
4
caddisflies (Trichoptera), and non-biting midges (Chironomidae). Prolonged and chronic
streamflow reductions can severely alter stream communities by stressing or eliminating flow-
dependent taxa. The absence of these taxa can, in turn, affect food webs and ecological
processes ecosystem-wide.
community structure used for characterizing invertebrate community quality include: species
as per Pielou 1977), diversity (as a combined measure of richness and evenness, e.g. Shannons
equation, Krebs 1999), relative abundance, and absolute abundance (no. of organisms per unit
surface area).
Fish communities and individual fish species have also been used as indicators of system
health and as a means of evaluating system disturbance (Karr 1981; Fausch et al. 1984; Miller et
al. 1988). Shifts in the relative abundance of obligate riverine, facultative riverine and
facultative lacustrine species may also provide indications of flow change. As previously
mentioned however, fishes tend to exhibit greater tolerance to flow changes than invertebrates,
particularly in peninsular Florida streams in which obligate riverine species are generally absent.
Thus, the effect of a natural flow deviation on fish populations is related to the sensitivity of
availability by fishes to identify thresholds and predict population and community level
responses. The fish species selected for this study are Suwannee bass Micropterus notius,
largemouth bass M. salmoides, American eel Anguilla rostrata and spotted sucker Minytrema
melanops. The Suwannee bass is a black bass that is generally confined to fluvial systems east
of the Ochlocknee River and west of the Suwannee River drainage. It is an important species for
5
recreational angling and is listed as a State of Florida Species of Special Concern (FAC 68A-
27.005). The largemouth bass, also a black bass, is highly valued by recreational anglers. It is
widely considered a habitat generalist and will serve to potentially elucidate habitat utilization
similarities or differences with Suwannee bass. The American eel is a species with little habitat
utilization information and was chosen due to its catadromous life history. The spotted sucker is
also predominantly a fluvial species often considered sensitive to poor water quality with an
OBJECTIVES
minimum flows and levels in their respective districts; however, few of these approaches
consider the biological communities that MFL legislation and policy were designed to include,
due in part to a lack of fish and wildlife information. The overall purpose of the research
presented herein was to begin development of ecology-based MFL information that would focus
upon protection of habitat and biological diversity and, thereby, provide fish and wildlife
guidance to the five water management districts. To achieve this purpose, the following
2. Develop habitat selection (preference) curves for selected fish and aquatic invertebrate
taxa sampled within the study area.
3. Model the relationships between hydrologic and habitat variables in selected stream
segments.
4. Using results from objectives one, two, and three, predict the usable habitat area for
selected fish and invertebrate species at variable flows and levels using an existing, or
developed, habitat simulation model.
6
These objectives were selected with the expectation that the ensuing product would
benefit the aquatic resources of Florida by providing support to the states water management
districts in the development of MFLs. The work will also benefit the Florida Comprehensive
Wildlife Strategy by addressing two required elements: (1) Descriptions of locations and relative
condition of key habitats and community types essential to conservation of declining species or
those indicative of ecosystem health, and (2) Descriptions of problems which may adversely
affect those species or their habitats. As such, the information gained is not necessarily limited
to the study stream, and may be transferable to other streams within the subject species
geographical ranges.
STUDY AREA
The study area was the northern Withlacoochee River (Madison and Hamilton Cos.,
Florida) from approximately 1 km upstream (north) of Florida State Route (SR) 6 highway to the
confluence with the Suwannee River near Ellaville, Florida (Figure 1, Table 1). The
Withlacoochee originates in south Georgia near the cities of Tifton and Moultrie and drains
approximately 6,035 km2 (2,330 mi2 ) of mostly agricultural lands in Georgia and Florida (Hand
et al. 1996). Most of the watershed lies in Georgia. The river flows through the Gulf Coastal
Lowlands Physiographic Zone and, in the study area, is underlain by Tertiary period
differentiated strata that include Ocala Limestone, Suwannee Limestone, the Torreya Formation,
the Statenville Formation, and the Miccosukee Formation (Scott et al. 1991). An extensive karst
system links the Florida aquifer to surface waters in the study area. A number of springs, most
prominently Madison County Blue Spring and Morgans Spring, arise along the river and supply
clear water of low dissolved oxygen and constant temperature to the stream. Madison County
Blue Spring is a first magnitude spring (annual median flow > 100 cfs) that discharges to the
7
FIGURE 1. The Withlacoochee River, Florida, study area ( = relative locations of stream
reaches containing transects surveyed for PHABSIM analysis).
8
TABLE 1. Geographic locations of Withlacoochee River, Florida, transects surveyed for
PHABSIM habitat simulation modeling.
9
Withlacoochee via a 50 meter spring run near Florida SR6 highway (Figure 1). Madison Blue
Spring supplies up to 20 percent of the total Withlacoochee discharge in the study area during
Within the study area the Withlacoochee is characterized by steep sand and limestone
banks and alternating sequences of deep pools, runs, limestone bedrock/cobble/gravel shoals, and
shifting sand shoals. Mean daily Withlacoochee River discharges during the study period ranged
from 245 cfs (1-02-08) to 17,900 cfs (3-02-08); Figure 2). The Withlacoochee crested above
flood stage after the passing of two hurricanes during the summer of 2004; however, the work
TECHNICAL APPROACH
The Physical Habitat Simulation Model (PHABSIM; Milhous et al. 1984, 1989), a
system of models for simulating available stream habitat, was selected as the most appropriate
tool for evaluating potential habitat loss resulting from reduced streamflows in the
Methodology (IFIM; Milhous et al. 1989, Stalnaker et al. 1995, Bovee et al. 1998), a prioritized
collection of field, modeling, and negotiation protocols intended to assist natural resources
managers and stakeholders in the determination and selection of appropriate stream management
weighted usable area (WUA). WUA quantifies the amount of micro-habitat available for a target
organism over a range of stream discharges. PHABSIM is the most commonly used simulation
model for determining potential impacts of reduced flows. It has been used in a variety of stream
10
FIGURE 2. Northern Withlacoochee River discharge at the Lee, Florida USGS streamflow
gauge (no. 02319394) during the gauge period of record (Nov. 2000 through Nov. 2008). The
FWC study was executed during the time period between the vertical red lines (1 July 2005 30
June 2008).
11
PHABSIM Transect Survey Methods
Transects were selected and surveyed using PHABSIM protocols as per Bovee (1997)
and Bovee et al. (1998). Study reaches and PHABSIM transects were selected during May 2006
with the assistance of Dr. James Gore of the University of South Florida. A total of 20 transects
were distributed among nine study reaches (Figure 1, Table 1). Transects were selected based
upon their flow characteristics (laminar, one dimensional), streambed morphology, presence of
microhabitats typical of the reach, and relation to instream hydraulic controls (mostly limestone,
bedrock shoals). Transects were marked by placing a permanent headpin at each transect start
point on the left bank, looking upstream, and a tail pin at each transect end on the right bank.
The pins were placed above the normal high water mark to allow location of transects in all
during both low flow and moderate flow conditions. The minimum number of measurement
increments (stations) along each transect was determined by dividing the bank-to-bank wetted
width of the river at the location by 20. The resulting quotient was the minimum number of
stations along the transect. Distance between stations never exceeded five feet (1.5 m). To
facilitate measurements a marked Kevlar tag line was placed perpendicular to stream flow at
each transect. The tag line was anchored to trees or augered tie-downs. Head and tail pins were
never used to tie-down the tag line. Habitat measurements were obtained by wading when water
depth was less than 1.5 m and stream flow conditions safely permitted. Measurements were
obtained from a boat using U.S. Geological Survey (USGS) boat equipment in depths greater
than 1.5 m or in high flow conditions. The USGS equipment was designed to stabilize a boat in
swift current. It consisted of an aluminum frame that was bolted to the boat gunnels and through
12
which a steel cable was threaded and affixed to each stream bank. The cable was stretched tight
with a come-along winch to stabilize the boat and allow movement perpendicular to flow.
Habitat measurements obtained at each station along each transect included distance from
headpin (interval- x), height of substrate above ground surface (i.e.; cobble or boulder), substrate
type (Table 2), water depth, and average current velocity. Water column depth was measured
using a 2 m pole calibrated in one centimeter increments when wading, or with a sounding reel
when using the boat. Current velocities were measured using a Marsh-McBirney Flo- mate
2000 portable meter. At depths of 100 cm or less, average current velocity was determined from
a measurement taken at 0.4 x total depth from the stream bottom (60 percent total depth
from surface; Smith and Stopp 1978). At depths exceeding 100 cm, average velocity was
obtained by taking the mean of measurements made at 0.2 and 0.8 x total depth (Smith and Stopp
1978). Bed elevation and water surface elevation were recorded using a total station surveying
instrument. Water surface elevations were measured along each transect at three locations: left
For each reach surveyed (whether a single transect or set of transects), a benchmark
was established to enable the linking of all distance and height measurements along the transect
to a common datum. Benchmarks were placed on permanent structures (trees, bedrock ledges)
above the normal high water mark. An instrument height was set at each benchmark to
establish a common datum for stream bed heights. Measurements of intervals, benchmarks, and
water surface elevations were recorded in 10th s of feet. Ground elevations and intervals were
measured to the nearest 0.1 ft. Water surface elevations were measured to the nearest 0.01 ft.
13
TABLE 2. Florida Fish and Wildlife Conservation Commission substrate
and cover type designations used for PHABSIM analysis of selected
northern Withlacoochee River fish and invertebrate taxa.
Cobble/gravel 3
Organic debris pack Includes fine woody debris and leaf litter 6
14
Low- flow measurements were obtained 8 June 6 July 2006, when discharge measured
at the USGS streamflow gauge at Lee, Florida, ranged between 460 cfs and 1,030 cfs. Moderate
flow measurements were obtained 4 February 19 February 2008, when discharges at the Lee
selected stream reaches (Figure 3) using habitat-specific quantitative methods. Habitats sampled
included limestone bedrock outcroppings, shallow cobble/gravel riffles, sand sediments, snags
(wood debris), tree root systems, and organic debris packs. One hundred eighteen samples were
obtained from eight study reaches (Figure 3) during the field component of the study.
Communities associated with limestone bedrock and cobble/gravel riffles were sampled
using a Hess sampler (Jacobi 1978, Merritt et al. 2008). Sand sediments in runs and pools were
sampled using a petite ponar dredge (Powers and Robertson 1967, Merritt et al. 2008). Snags
(wood debris) were sampled using a bag and saw technique developed by Warren et al. (2000).
Tree root systems and benthic organic debris packs were sampled using a quantitative dip-net
technique developed by Warren et al. (2006). All sampling and rinsing devices were fitted with
300 mesh Nitex to enable collection of smaller-sized invertebrates. Upon collection, each
sample was rinsed in a 300 mesh sieve bucket to remove excess water, then preserved
each individual sample. Bottom substrate type was visually characterized according to the
modified Wentworth scale. Average water column current velocity and nose depth current
velocity were measured with a Marsh-McBirney Flo-Mate Model 2000 portable meter. Bottom
15
N
SR 6
Reach 0
Reach 3
Withlacoochee River
Reach 3.5
Reach 3.75
141
Morgan Spring CR
Reach 6
US 90
0 Kilometers 3 Reach 7
Suwannee River
FIGURE 3. Relative locations of Florida Fish and Wildlife Conservation Commission aquatic
invertebrate sampling sites in the northern Withlacoochee River, Florida, study area, 2005-2007.
16
depth was measured with a meter stick or calibrated wading rod. Water temperature, dissolved
oxygen, and specific conductance were measured using a YSI Model 556 multi-parameter meter
at the surface, bottom, average water column depth and nose depth.
Samples were transported to the Florida Fish and Wildlife Conservation Commission
Freshwater Invertebrate Laboratory in Gainesville for processing. Each sample was processed
separately; samples were not composited. Processing was conducted by trained laboratory
small sample portions in water-filled glass petri plates and then, with the aid of forceps and a
stereo-scope, removed invertebrates. The technicians identified and sorted invertebrates to major
taxonomic groups, enumerated each taxon, and preserved the organisms with 95 percent ethanol
in three-dram vials. A taxonomist then verified counts and identified all organisms in each
sample to the lowest taxonomic level possible, given the age and physical condition of
Simuliidae, specimens of these taxa were slide- mounted using CMC-10 and then identified using
A separate laboratory sheet was prepared for each sample. Taxonomic composition and
taxa abundances were recorded on these sheets, along with ancillary measurement data, and
station/date identifiers. Sample site information, ancillary measurements, taxa codes, and raw
organism counts were entered from laboratory sheets into a database maintained in Microsoft
Excel. Taxa richness, diversity (Shannons equation as per Krebs 1999), and evenness (Pielou
1977) were computed for each sample and entered into the database. Descriptive statistics and
univariate analyses were computed using SYSTAT 12 (SYSTAT Software, Inc. 2007) and
JMP 7 (SAS Institute Inc., 2007) statistical software. Multivariate analyses were computed
17
using PC-ORD Version 5 (McCune and Mefford 1999). Habitat preference curves for selected
invertebrate indicator taxa were developed for input into the PHABSIM stream habitat
simulation model by Dr. James Gore of the University of South Florida at St. Petersburg using
Withlacoochee River data and the methods of Gore and Judy (1981). Dr. Gore executed the
PHABSIM model using the selected taxa and computed time series analyses to determine
weighted usable habitat areas available at study transects throughout the entire discharge range
for the period of record at the USGS Pinetta stream gauge on the Withlacoochee River.
We used a 4.9 m aluminum electrofishing boat equipped with a Smith-Root 5.0 GPP to
collect juvenile and adult largemouth and Suwannee basses, juvenile and adult spotted suckers,
and American eels within the study area depicted in Figure 1. Because the targeted species tend
to associate with instream structure, 200 m long transects were located parallel to the riparian
zone, where the majority of instream cover existed in the form of overhanging terrestrial
vegetation, tree roots, limerock outcroppings and within-channel snags. Sampling was
and fall (October-December) months to account for temporal shifts in habitat utilization.
Transects were predominantly located within run mesohabitats and effort was made to plan
collection trips near baseflow discharges during each season. Thus, river bends and riffle
habitats were generally avoided where water depth and clarity limited the electrofishing
efficiency, and where water velocity was too great to effectively capture stunned fishes,
respectively. Similarly, sampling near base flow conditions minimized stunned- fish drift during
electrofishing and maximized water clarity to more accurately determine the fishs location and
18
Prior to fish sampling, 5 numbered buoys were randomly deployed within each 200 m
transect for available habitat estimation. During fish sampling, a numbered buoy was deployed
over the habitat where each target individual was encountered for utilized habitat estimation.
Buoys were not deployed if the original location of the target species was uncertain. Collected
target fish were measured to total length (millimeters) and returned to the water. Subsequent to
fish sampling, microhabitat measurements were recorded at each buoy corresponding to either
available or utilized habitats. Water velocity (m/s) was recorded at 0.6 of the depth in the water
column using a Marsh-McBirney Flo- mate 2000 portable velocity meter. Water column depth
(cm) was recorded using a 2 m pole in centimeter increments, or a Speedtech Depthmate model
SM-5 if the depth exceeded 2 m. Cover and substrate type were qualitatively visually described
and were assigned to categories (Table 2). Both water velocity and depth were recorded at the
buoy, whereas cover and substrate was described within a 0.91 m radius of the buoy.
Length frequency histograms were constructed for largemouth and Suwannee basses and
spotted suckers from each sampling event to identify the length break between juvenile and adult
fishes. We assumed that the first mode in the distribution represented juveniles as either age-0 in
the summer and fall collections, or age-1 fishes when collected the winter and spring. All
The data were separated by sampling event, species, and life stage to examine if habitat
utilization relative to habitat availability within seasons served as reasonable replicates. Habitat
suitability criteria per species were developed using category III criteria methods described by
Bovee (1986). The data were organized into bins of 6 cm/s water velocity intervals, 30 cm depth
intervals, and cover/substrate categories shown in Table 2. The suitability (selection) of velocity,
19
depth and cover to target species was demonstrated by independently calculating the ratio
between the relative frequencies of utilization and availability across the variable intervals using
Si = Ui / Ai,
where Si is an index of suitability at xi, Ui is the relative frequency of fish observations at xi, Ai is
the relative frequency of xi available during the sampling events, and xi is the interval of the
variable x. The suitability index was then standardized to provide a scale from 0 to 1 by dividing
the ratios across the variable intervals by the maximum ratio value. Ratios must be standardized
for use with PHABSIM since the maximum value of any weighting factor used in the program is
one (Bovee 1986). Because the suitability equation is a function of both habitat utilization and
habitat availability, the term selection is often used interchangeably with suitability throughout
the remainder of the fish portion of this document. However, we offer no defining value or
threshold to discern habitat selection verses avoidance. Rather, statements regarding selection or
suitability are quantified within the scale of zero to one, but are primarily used to express the
RESULTS
groups were collected from the eight river reaches sampled within the study area (Appendix 1).
The overall river assemblage was numerically predominated by larval non-biting midges
20
Community structure varied substantially among the micro-habitats sampled (although some taxa
were ubiquitously distributed among all habitats). The most species rich, diverse, and densely
populated communities were present in micro-habitats that were characterized by greater current
velocities and higher levels of dissolved oxygen (bedrock and cobble /gravel riffles), and in
micro- habitats that were located above bottom sediments (snags and tree root systems), but not
necessarily in higher current velocities. Bottom-associated habitats (sand and organic debris
The invertebrate assemblage associated with snags (above-bottom wood debris) was the
most species rich (133 total taxa) and diverse (mean diversity = 3.77) of all communities
sampled (Table 3); however, snags ranked fourth among the six habitats sampled in terms of
mean total organisms per square meter (29,413 m-2 ; Table 4). The snag-associated community
was predominated by the non-biting midges Tanytarsus species C (3,253 m-2 , 11.1%),
Pseudochironomus sp. (3,068 m-2 , 10.4%), and Xestochironomus sp. (1,520 m-2 , 5.2%), the riffle
beetle Stenelmis sp. (2,904 m-2 , 9.9%), and water mites (Hydracarina (1,693 m-2 , 5.8%;
Appendix 2D). These five taxa together accounted for 42.4 percent of all organisms collected
from snags. No other individual taxon accounted for more than 4.4 percent of the total snag
organisms.
The invertebrate community associated with tree root systems was nearly equivalent to
the snag-associated community in terms of total taxa richness, mean taxa richness, and Shannon
diversity, however, taxonomic composition of the two habitats differed substantially. One
hundred twenty-three total taxa were collected from tree roots. Per sample values of species
richness and Shannon diversity averaged 35.0 and 3.77, respectively (Table 3). The non-biting
midge Tanytarsus sp. C was by far the most abundant root-associated taxon (9,324 m-2 ),
21
TABLE 3. Descriptors of invertebrate community quality in ecologically important habitats in
the northern Withlacoochee River, Florida, 2005-2007: total taxa richness, and per sample
means of taxa richness, diversity (Shannons equation), and evenness (as per Pielou 19xx).
Habitat _
Bedrock Cobble Riffle Sand Snags Tree Roots Debris Packs
22
TABLE 4. Density (no. m-2 ), coefficient of variation (c.v.), and percent composition (%) of key invertebrate indicator taxa collected
from six ecologically important habitats in the northern Withlacoochee River study area, Florida, 2005-2007.
Habitat _
Bedrock _ Cobble Riffle Sand _ Snags _ Tree Roots_ Debris Packs
no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.)
Taxon % % % % % %
Gastropoda (total) 107 (1.62) 835 (1.36) 246 (2.07) 92 (1.75) 1,679 (1.09) 48 (0.35)
0.3 1.5 1.8 0.3 2.8 0.3
Amphipoda (total) 71 (1.60) 173 (1.78) 42 (2.01) 966 (2.61) 5,627 (1.17) 97 (0.69)
0.2 0.3 0.3 3.3 9.4 0.5
Hyalella azteca 33 (1.74) 103 (1.55) 34 (2.22) 903 (2.64) 2,196 (1.80) ---
<0.1 0.2 0.2 3.1 3.7
Aquatic Acari (Hydracarina) 659 (0.71) 5,600 (1.17) 268 (1.66) 1,693 (0.74) 1,341 (1.05) 280 (0.96)
1.7 10.1 2.0 5.8 2.2 1.6
Ephemeroptera (total) 788 (1.19) 3,104 (1.03) 48 (2.45) 1,215 (0.80) 2,596 (1.01) 261 (0.62)
2.0 5.6 0.4 4.1 4.3 1.5
Total Baetidae 176 (1.91) 484 (1.06) 12 (2.70) 170 (1.23) 208 (1.17) 87 (1.73)
0.4 0.9 <0.1 0.6 0.3 0.5
23
TABLE 4 (cont.)
Habitat _
Bedrock _ Cobble Riffle Sand _ Snags _ Tree Roots_ Debris Packs
no. m-2 (c.v.) no. m-2 (c.v.) -2
no. m (c.v.) no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.)
Taxon % % % % % %
Megaloptera (hellgrammites)
Corydalus cornutus 5 (2.90) 105 (2.40) --- 7 (2.01) --- 5 (1.73)
<0.1 0.2 <0.1 <0.1
Trichoptera (total) 2,000 (0.94) 2,916 (0.95) 165 (3.18) 3,691 (1.16) 7,542 (0.77) 431 (0.37)
5.1 5.3 1.2 12.5 12.6 2.4
Total Hydropsychidae 323 (1.31) 461 (1.84) 14 (4.30) 968 (1.30) 63 (3.52) 29 (1.31)
0.8 0.8 0.1 3.3 0.1 0.2
Hydropsyche rossi 213 (1.33) 137 (1.70) 1 (6.24) 528 (1.71) 7 (3.87) 5 (1.73)
0.5 0.2 <0.1 1.8 <0.1 <0.1
Total Hydroptilidae 1,386 (1.02) 2,061 (1.01) 74 (3.21) 1,291 (0.90) 4,422 (0.91) 193 (0.96)
3.4 3.7 0.5 4.4 7.4 1.1
Hydroptila sp. 1,192 (1.06) 1,566 (1.12) 49 (3.42) 1,000 (1.15) 1,748 (0.83) 39 (0.87)
3.1 2.8 0.4 3.4 2.9 0.2
Coleoptera (beetles)
Stenelmis sp. (larvae) 4,536 (1.77) 8,783 (2.08) 162 (1.36) 2,904 (0.76) 2,299 (1.24) 358 (0.17)
11.7 15.9 1.2 9.9 3.8 2.0
24
TABLE 4 (cont.)
Habitat _
Bedrock _ Cobble Riffle Sand _ Snags _ Tree Roots_ Debris Packs
no. m-2 (c.v.) no. m-2 (c.v.) -2
no. m (c.v.) no. m-2 (c.v.) no. m-2 (c.v.) no. m-2 (c.v.)
Taxon % % % % % %
Total Chironomidae (midges) 22,137 (0.95) 28,049 (1.05) 6,447 (1.04) 15,395 (0.73) 31,720 (0.57) 15,167 (0.58)
57.0 50.8 47.2 52.3 53.0 85.8
Cricotopus bicinctus 688 (2.44) 853 (1.80) 33 (4.73) 228 (1.92) 4,673 (1.05) 174 (1.45)
1.8 1.5 0.2 0.8 7.8 1.0
Parakiefferiella sp. 2,259 (2.15) 1,131 (0.88) 254 (3.42) 1,281 (2.19) 2,805 (3.87) 522 (0.89)
5.8 2.0 1.9 4.4 4.7 3.0
Tvetenia vitracies 1,229 (2.02) 393 (2.07) 113 (3.99) 983 (2.73) 67 (2.45) 58 (1.73)
3.2 0.7 0.8 3.3 0.1 0.3
Pseudochironomus sp. 7,537 (1.37) 4,965 (1.49) 476 (3.50) 3,068 (1.92) 296 (1.92) 135 (0.89)
19.4 9.0 3.5 10.4 0.5 0.8
Stempellina sp. A 122 (3.61) 2,073 (2.71) 855 (1.78) 72 (2.03) 542 (2.06) 19 (1.73)
0.3 3.8 6.3 0.2 0.9 0.1
Tanytarsus sp. C 3,362 (1.61) 11,942 (1.41) 465 (2.14) 3,252 (1.63) 9,324 (1.32) 232 (1.00)
8.6 21.6 3.4 11.1 15.6 1.3
Total Organisms 38,865 (0.67) 55,241 (0.88) 13,665 (0.84) 29,413 (0.50) 59,802(0.59) 17,686 (0.47)
100.0 100.0 100.0 100.0 100.0 100.0
25
accounting for 15.6 percent of all organisms collected (Appendix 2 E). No other individual taxon
accounted for more than 4.9 percent of the total. Other numerically important root-associated
taxa included the amphipods Gammarus sp. (2,905 m-2 , 4.9%) and Hyalella azteca (2,196 m-2 ,
3.7%), the riffle beetle Stenelmis sp. (2,299 m-2 , 3.8%), and the midge Parakiefferiella sp. (1,870
m-2 , 3.1%; Appendix 2E). Mean total organisms in tree roots (59,802 m-2 ) was the highest of all
The bedrock and cobble/gravel riffle habitats supported high quality invertebrate
communities that were slightly subordinate to the snag and tree root-associated communities in
terms of taxa richness and diversity. Ninety and 115 total taxa were obtained from the bedrock
and cobble/gravel habitats, respectively (Table 3). Mean taxa per sample averaged 27.3 and 38.3
in bedrock and cobble/gravel, respectively, while Shannon diversity averaged 3.07 and 3.49,
respectively (Table 3). Tanytarsus sp. C, Stenelmis sp., and Pseudochironomus sp. were the
three most abundant taxa in both the bedrock and cobble/gravel habitats. Together these three
taxa accounted for 46.5 percent of all organisms collected from cobble/gravel and 39.7 percent of
The sand and organic debris pack-associated invertebrate communities exhibited the
lowest mean diversities (2.75 and 2.97, respectively) and lowest mean total organism densities
(13,665 m-2 and 17,686 m-2 , respectively) of all Withlacoochee River habitats sampled (Tables 3
and 4). Both habitats were numerically predominated by a single taxon that was far more
abundant than any other single organism, thus contributing to the lower mean diversity values.
The Asian clam Corbicula fluminea was the most abundant taxon collected from sand (2,042 m-
2
), accounting for 14.9 percent of the total organisms (Appendix 2C). The lone remaining
organism accounting for greater than four percent of the total sand organisms was the non-biting
26
midge Stempellina sp. A (Appendix 2C). The orthocladine midge Corynoneura sp. B accounted
for 50.2 percent (8,884 m-2 ) of all organisms collected from organic debris packs (Appendix 2F).
Midges of the genus Corynoneura are thought to be pollution-sensitive (Simpson and Bode
1980). The only remaining debris pack-associated taxon that occurred in densities greater than
1,000 m-2 and accounted for more than three percent of the total was the nonbiting midge
Functional feeding group analysis (Cummins 1973 and 1974, Hawkins and Sedell 1981,
Merritt et al. 2002) was conducted upon habitat-specific invertebrate assemblages to evaluate the
Withlacoochee River in the study area in terms of the nutritional resource base available and to
provide a baseline for future analyses focused upon potential stream flow reduction impacts and
associated consequences to food resource availability and the food web. Invertebrate taxa were
assigned to functional feeding groups according to the determinations of Barbour et al. (1999)
Results of the analysis indicated that all sampled habitats were numerically dominated by
the gathering and filter feeding functional groups, which, together accounted for between 53.6
percent (tree roots) and 75.6 percent (debris packs) of the total organisms in each habitat (Figure
4). Gatherers were the single most abundant group in every habitat sampled, while filterers were
second- most abundant in all habitats except organic debris packs, where they constituted only
4.0 percent of the total. Both gatherers and filter feeders utilize upon fine particle size (< 1 mm)
organic matter (FPOM; primarily detritus) as a food resource (Cummins et al. 2008). Filterers
are especially important in the context of flow because many (e.g. net spinning caddisflies) rely
upon flow to inflate spun nets and to provide a drifting source of FPOM. Periphyton scrapers
accounted for between 17.1 and 22.5 percent of the total fauna in habitats characterized by
27
TABLE 5. Florida Fish and Wildlife Conservation Commission aquatic invertebrate
functional feeding group designations (based upon and modified from Barbour et al.
1999 and Cummins et al. 2008).
28
100%
90%
80%
70%
60%
Shredders
Scrapers
50% Gatherers
Filterers
40% Predators
Generalists
30% Parasites
20%
10%
0%
Bedrock Cobble Sand Snags Tree Roots Debris Packs
Habitat
FIGURE 4. Aquatic invertebrate functional feeding group composition within the six Withlacoochee River habitats sampled during
2005 2007.
29
harder substrates (bedrock, cobble, snags, and tree roots), and accounted for less than 6 percent
of the total in the softer sand and debris pack habitats (Figure 4). The predator functional group
was a prominent component in the cobble, sand, and snag-associated communities (12.4, 11.4,
11.9 %, respectively), but accounted for less than 7.5 percent of the total abundance in bedrock,
tree roots, and debris packs. The generalist functional group (primarily omnivores and
scavengers) was prominent only in the organic debris pack habitat, where it was the second-most
abundant group collected and accounted for 13.6 percent of the fauna (Figure 4). Shredders did
not account for more than 3.6 percent of the total organisms in any habitat (Figure 4),
demonstrating the general lack of coarse particulate organic matter (CPOM) in the study area.
Habitat suitability criteria constitute the primary biological input to the PHABSIM
model. The product of PHABSIM, weighted usable habitat area, is a func tion of the suitability
of current velocity, depth, and substrate for a given taxon or community quality descriptor at a
given stream transect/discharge. Suitability curves containing the biological data for input into
PHABSIM can be developed from data derived using a number of methods ranging from best
(current velocity, depth, and substrate) and target organism abundances across ranges of the
habitat parameters (Bovee 1986, Gore and Judy 1981). To construct the curves, suitability for
the current velocity, depth, and substrate parameters is relativized on a scale ranging from zero to
one, with zero representing the least optimum value and one representing the habitat value at
which populations of the subject taxon/community quality parameter attain optimum levels.
Suitability curve construction using data collected on-site allows the establishment of both a
usable range and an optimum range of a given habitat variable for a given species or descriptor
30
in the study stream (Bovee 1986, Gore 1987, Gore et al. 2001). The authors and users of the
PHABSIM model consider the optimum range of a given variable to be 0.85 and above on the
greater than 15 percent from the optimal level is considered to be ecologically harmful.
abundance and community quality parameter data collected directly from the Withlacoochee
River study area. Current velocity, depth, and substrate type were measured with each individual
invertebrate sample collected. Using these data, habitat suitability curves for selected
Withlacoochee River community quality parameters and invertebrate taxa were developed by Dr.
James Gore of the University of South Florida at St. Petersburg, following the methods of Gore
and Judy (1981). Community quality parameters and individual invertebrate taxa for which
curves were developed were selected based upon criteria that included flow dependence,
abundance across all sampled habitats, and importance as a community quality indicator. The
community quality parameters and taxa for which curves were developed are presented in Table
6. All invertebrate suitability curves produced are contained in Appendix 3. Example habitat
suitability curves for net-spinning caddisfly larvae of the genus Hydropsyche and larvae of the
baetid mayfly Pseudocloeon ephippiatum are presented in Figures 5 and 6. The optimum ranges
of flow and depth for all selected indicator taxa and community quality descriptors are presented
in Table 7.
invertebrate taxa that are dependent upon flow for survival. Hydropsyche spp. and other net-
(Appendix 1) require flow to provide food resources and to inflate capture nets (Edington 1968,
31
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
FIGURE 5. Current velocity, depth, and substrate suitability curves for the net-spinning
caddisfly Hydropsyche sp. (Trichoptera:Hydropsychidae) collected in the northern
Withlacoochee River, Florida, 2005-2007.
32
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
FIGURE 6. Current velocity, depth, and substrate suitability curves for the mayfly
Pseudocloeon ephippiatum (Ephemeroptera:Baetidae) collected from the northern
Withlacoochee River, Florida, 2005-2007.
33
TABLE 6. Aquatic invertebrate community descriptors and taxa selected for habitat
suitability curve determination and PHABSIM analysis.
34
TABLE 7. Optimum ranges of current velocity and depth for the selected community
quality parameters and indicator invertebrate taxa as determined by habitat suitability
curve development.
Optimum Ranges _
Community Descriptor/Taxon Current Vel. (cm/sec.) Depth (cm)
35
Wallace 1975). Pseudocloeon ephippiatum occurs on plants and hard substrates in the swiftest
areas of streams where it grazes algae, diatoms, and biofilm, and depends upon flow to
oxygenate gills (Berner and Pescador 1988 as Baetis ephippiatus). Habitat suitability curves
cm/sec. and an optimum depth range of 61 to 114 cm. The Hydropsyche substrate suitability plot
demonstrates a preference for the snag habitat (Figure 5). Habitat suitab ility curves for
and a depth range of 69 to 122 cm would support an optimum population level. In contrast to
Examination of current velocity suitability curves for the community quality descriptors
taxa richness, diversity, and total organisms indicated that a range of 37 to 60 cm/sec would
support optimum values for those parameters (Table 7). Optimum depths for the three
descriptors ranged from 61 to 105 cm (Table 7). Optimum current velocity ranges for individual
taxa ranged from a low of 0 3 cm/sec for the chironomid Stempellina sp. A to a high of 79
183 cm/sec for the chironomid Parakiefferiella sp. Optimum velocities for the flow dependent
cornutus) taxa were moderate among the individual taxa and ranged from 34 to 86 cm/sec.
Optimum depth ranges for individual taxa ranged from a low of 9 22 cm for Stempellina sp. A
to a high of 110 to 139 cm for the chironomid Pseudochironomus sp. Optimum depths for the
flow dependent mayflies, caddisflies, and hellgrammites again ra nged among the moderate
36
Invertebrate Community PHABSIM Analysis
The PHABSIM model was used to determine areas of usable habitat (WUAs) for selected
invertebrate indicator taxa and community quality d escriptors (Table 6) at given discharges
within the eight PHABSIM study reaches (Table 1). Graphic representations of PHABSIM
results for all taxa and descriptors are presented in Appendix 4. Optimum flows (those that
produced the greatest usable habitat areas) for mean (per sample) taxa richness ranged from 135
cfs at Reach 6 to 650 cfs at Reach 1 (Table 8). Optimum community diversity was attained at
flows ranging from 280 cfs at Reach 5 to 650 cfs at Reaches 1 and 2 (Table 8). The greatest
number of total organisms per sample occurred at discharges ranging from 700 cfs at Reach 7 to
2,000 cfs at Reach 5 (Table 8). Optimum flows for individual taxa ranged from a low of 135 cfs
for both the riffle beetle Stenelmis sp and the midge Stempellina sp. A at Reach 5 to a high 2,500
To conduct time series analyses that would utilize Withlacoochee River streamflow data
encompassing the longest possible time period, we regressed mean monthly discharges at the
Lee, Florida, streamflow gage (period of record = Nov. 2000 - present) against mean monthly
discharges measured at the Pinetta, Florida gage (period of record = Oct. 1931 - present) for the
Lee gage period of record (R2 =0.95, p<0.05). The Lee gage is located within the Withlacoochee
study area between Reaches 6 and 7. Using the regression equation, monthly values for the Lee
gage were then computed from October 1931 to the beginning of the Lee gage period of record.
Results of time series analyses showed that, in general, flow reductions during the typical
higher flow months of January through April produce increases in habitat area (WUA) at all
study reaches (Tables 9 - 11, Appendix 5). However, flow reductions during the months of May
37
TABLE 8. Discharge (Q, in cubic feet per second [cfs]) at which optimum weighted usable area (WUA, in sq. ft. of habitat/1000 linear feet of
stream) is attained for community quality descriptors and individual indicator invertebrate taxa at the seven PHABSIM study reaches in the
northern Withlacoochee River, Florida.
Stream Reach _
Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Q Q Q Q Q Q Q Q
Taxon (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA)
Taxa Richness 650 600 400 280 340 300 135 220
(124,863) (100,743) (40,467) (29,627) (96,474) (88,002) (31,531) (79,985)
Shannon Diversity 650 650 400 400 340 280 400 400
(131,785) (98,278) (37,714) (24,099) (102,721) (94,517) (26,657) (71,657)
Total Organisms 1,200 1,000 950 1,100 1,500 2,000 900 700
(89,595) (82,157) (30,165) (10,159) (68,851) (82,287) (12,462) (36,517)
Total Baetidae 1,300 1,100 1,150 1,500 2,000 750 1,200 950
(51,742) (62,169) (24,152) (2,247) (34,005) (46,344) (4,992) (12,988)
Pseudocloeon ephippiatum 650 575 650 900 1,050 340 700 575
(124,863) (41,825) (19,221) (854) (15,680) (28,129) (1,463) (3,219)
Total Hydropsychidae 1,200 900 750 475 1,200 340 700 340
(34,891) (31,942) (8,314) (927) (23,055) (14,602) (2,904) (8,974)
38
TABLE 8 (cont.).
Stream Reach _
Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Q Q Q Q Q Q Q Q
Taxon (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA)
Hydropsyche sp. 1,150 1,050 950 200 1,200 650 750 340
(35,132) (27,298) (5,903) (799) (24,549) (11,370) (3,355) (9,292)
Hydroptila sp. 1,100 950 900 1,150 1,300 700 900 750
(74,734) (63,163) (22,281) (2,955) (53,194) (47,293) (8,230) (21,685)
Corydalus cornutus 750 525 650 900 950 340 650 575
(18,404) (39,032) (16,812) (610) (9,387) (25,260) (368) (988)
Stenelmis sp. 220 475 250 135 135 200 135 160
(55,825) (50,888) (25,956) (9,384) (39,353) (64,447) (9,386) (16,389)
Total Chironomidae 1,500 1,300 1,300 1,200 2,000 800 1,300 950
(97,750) (86,267) (30,575) (12,659) (68,961) (63,709) (13,907) (44,665)
Cricotopus bicinctus 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500
(18,432) (18,768) (11,688) (12,353) (16,179) (28,932) (6,663) (11,204)
Parakiefferiella sp. 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,000
(81,509) (59,451) (19,203) (10,166) (69,824) (72,432) (16,084) (39,460)
Tvetenia vitracies 1,150 1,200 1,000 1,050 1,300 1,150 850 700
(63,065) (33,040) (8,214) (2,156) (43,982) (25,090) (7,966) (21,424)
39
TABLE 8 (cont.).
Stream Reach _
Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Q Q Q Q Q Q Q Q
Taxon (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA) (WUA)
Pseudochironomus sp. 1,500 1,500 1,500 2,000 2,500 2,500 1,300 1,300
(67,711) (30,106) (13,423) (1,781) (34,459) (43,177) (7,682) (25,738)
Stempellina sp. A 190 200 135 135 135 190 135 250
(23,216) (10,165) (7,186) (7,841) (2,784) (7,639) (3,571) (7,454)
Tanytarsus sp. C 1,300 1,050 1,100 2,500 2,000 2,500 1,150 850
(47,789) (63,848) (26,120) (5,132) (34,058) (70,278) (4,823) (12,997)
40
TABLE 9. Time series analysis of PHABSIM results: months of habitat loss exceeding 15 percent at 20 percent flow reductions in the
northern Withlacoochee River, Florida, PHABSIM study reaches.
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Taxa Richness --- --- --- --- --- --- --- ---
Shannon Diversity --- --- --- --- --- --- --- ---
Total Organisms --- Oct-Nov --- --- --- --- --- ---
Total Baetidae --- Oct-Nov --- --- --- --- --- ---
Pseudocloeon ephippiatum --- --- --- Oct-Nov --- --- --- ---
Total Hydropsychidae --- Oct-Nov --- --- --- --- --- ---
Hydropsyche spp. --- Oct-Dec --- --- --- --- --- ---
Hydroptila sp. --- --- --- --- --- --- --- ---
Corydalus cornutus --- --- --- Sep-Jan --- --- --- ---
Stenelmis sp. --- --- --- --- --- --- --- ---
Total Chironomidae Nov Oct-Nov --- --- --- --- --- ---
Cricotopus bicinctus --- --- --- --- --- --- --- ---
41
TABLE 9 (cont.).
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Parakiefferiella sp. --- Aug-Nov --- --- --- --- --- ---
Tvetenia vitracies Oct-Nov Oct-Nov --- --- --- --- --- ---
Pseudochironomus sp. Jun-Dec Jun-Dec Jun-Dec Jun-Nov Jun-Dec Jun-Nov Aug-Nov Aug-Dec
Stempellina sp. A --- --- --- --- --- --- --- ---
Tanytarsus sp. C --- --- --- --- --- --- --- ---
42
TABLE 10. Time series analysis of PHABSIM results: months of habitat loss exceeding 15 percent at 30 percent flow reductions in
the northern Withlacoochee River, Florida, PHABSIM study reaches.
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Taxa Richness --- --- --- --- --- --- --- ---
Shannon Diversity --- --- --- --- --- --- --- ---
Total Organisms Oct-Nov Oct-Nov --- --- --- --- --- ---
Total Baetidae Sep-Dec Oct-Nov Oct-Nov Oct-Nov --- --- Oct-Nov Nov
Pseudocloeon ephippiatum Nov --- --- Sep-Dec --- --- --- ---
Total Hydropsychidae Oct-Nov Oct-Nov --- --- --- --- --- ---
Hydropsyche spp. Oct-Nov Oct-Dec --- --- --- --- --- ---
Hydroptila sp. Oct-Nov --- --- Oct-Nov --- --- --- ---
Corydalus cornutus Nov --- --- Aug-Jan --- --- Nov ---
Stenelmis sp. --- --- --- --- --- --- --- ---
Total Chironomidae Jun-Dec Oct-Nov Oct-Nov Oct-Nov --- --- Nov ---
Cricotopus bicinctus Jan-Apr --- --- --- --- --- --- ---
43
TABLE 10 (cont.).
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Parakiefferiella sp. Sep-Nov Aug-Nov Nov --- --- --- Nov Nov
Tvetenia vitracies Sep-Dec Oct-Nov Oct-Nov --- --- --- Nov ---
Pseudochironomus sp. May-Dec Jun-Dec Jun-Dec Jun-Dec Jun-Dec May- Dec Jun-Dec Jun-Dec
Stempellina sp. A --- --- --- --- --- --- --- ---
Tanytarsus sp. C Oct-Nov --- --- Sep-Nov --- --- --- ---
44
TABLE 11. Time series analysis of PHABSIM results: months of habitat loss exceeding 15 percent at 40 percent flow reductions in
the northern Withlacoochee River, Florida, PHABSIM study reaches.
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Taxa Richness --- --- --- --- --- --- --- ---
Shannon Diversity --- --- --- --- --- --- --- ---
Total Organisms SepDec Oct-Nov Oct-Nov Nov --- --- Nov ---
Total Baetidae Sep-Dec Oct-Dec Sep-Dec Sep-Dec Nov --- Sep-Dec Oct-Dec
Pseudocloeon ephippiatum Oct-Nov --- --- Sep-Dec --- --- --- ---
Total Hydropsychidae Sep-Dec Oct-Dec Nov --- --- --- --- ---
Hydropsyche spp. Sep-Dec Sep-Dec Oct-Dec --- --- --- --- ---
Hydroptila sp. Sep-Dec Oct-Dec Oct-Dec Sep-Nov --- --- Oct-Dec ---
Corydalus cornutus Nov --- Nov-Dec May-Dec --- --- Oct-Dec ---
Stenelmis sp. --- --- --- --- --- --- --- ---
Total Chironomidae Jun-Dec Aug-Dec Sep-Dec Sep-Dec Nov --- Sep-Dec Nov-Dec
Cricotopus bicinctus Dec-Aug Sep-Nov Jun-Sep --- --- --- --- Sep&Nov
45
TABLE 11 (cont.).
Location _
Descriptor/Indicator Taxon Reach 1 Reach 2 Reach 3 Reach 3.5 Reach 4 Reach 5 Reach 6 Reach 7
Parakiefferiella sp. Jun-Dec May-Dec Jun-Nov Aug-Nov Jun-Nov Jun-Nov Aug-Nov Sep-Nov
Tvetenia vitracies Sep-Dec Aug-Dec Sep-Dec Nov --- --- Oct-Dec Nov
Pseudochironomus sp. May-Dec May-Dec May-Dec May-Dec May-Jan May-Dec May-Dec May-Dec
Stempellina sp. A --- --- --- --- --- --- --- ---
Tanytarsus sp. C Sep-Nov Oct-Nov Oct-Nov Jun-Dec --- --- Oct-Nov ---
46
through December, historically the lower flow months, produce loss of habitat area for some
indicator taxa at some transects. Some flow reductions result in losses of habitat area that exceed
the critical 15 percent level for some taxa (Tables 9 - 11, Appendix 5).
The time series analysis for the net-spinning caddisfly Hydropsyche sp. at study Reach 1
is characteristic of the typical Withlacoochee River pattern (Figure 7). Flow reductions of up to
40 percent during January through May result in gains of habitat area exceeding 80 percent
(February, Table 8). Conversely, flow reductions of up to 40 percent in the autumn months
Time series analysis for the chironomid Pseudochironomus sp. at Shoal 5 illustrates the
most extreme impacts of flow reduction in the summer and autumn months (Figure 8). Habitat
area at Shoal 5 increases with flow reductions during the winter months of February and March;
however, beginning in May, flow reductions of up to 40 percent result in decreases in the area of
It is important to note that flow reductions of as much as 40 percent did not result in
significant habitat area losses ( 15 percent) for the community quality descriptors Shannon
diversity and mean taxa richness at any study reach (Tables 9 - 11, Appendix 5).
transects per event (Table 12). A fish sample was not collected in the fall of 2006. Over the
course of the study, we made 603 available habitat measurements, and collected 766 largemouth
bass, 647 Suwannee bass, 514 spotted suckers and 108 American eels. Microhabitat utilization
sampling points collected for spotted suckers and black basses were segregated according to
juvenile or adult life stage based on length frequencies within each sampling event (Table 12).
47
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
April
August
July
February
January
December
September
June
October
May
November
March
FIGURE 7. Time series comparison of flow reduction effects upon usable habitat area for Hydropsyche sp.
(Trichoptera:Hydropsychidae) at Withlacoochee River Shoal 1
48
20%
10%
0%
-10%
Habitat Gain / Loss
-40%
-50%
-60%
-70%
April
August
July
December
January
February
September
June
November
May
October
March
FIGURE 8. Times series comparison of flow reduction effects upon usable habitat area for Pseudochironomus sp.
(Diptera:Chironomidae) at Withlacoochee River Reach 5.
49
Table 12. The number of electrofishing transects, the number of available microhabitat sampling
points, and the number of utilized microhabitat (MH) sampling points (per life stage) during each
seasonal collection on the Withlacoochee River, FL. AMEE = American eel, LMB = largemouth
bass, SUBA = Suwannee bass, and SPSK = spotted sucker.
50
The 2005 year class of largemouth bass was absent relative to the 2006 year class, represented by
0 juveniles in fall 2005, 2 juveniles in winter 2006 and 8 juveniles in spring 2006. Similarly, the
2005 year classes of Suwannee bass and spotted suckers were absent with 4, 0 and 0 juveniles
and 0, 0 and 0 juveniles collected respectively, in the fall 2005, winter 2006 and spring 2006
sampling events. It also appeared that the 2007 year class of spotted suckers was absent with no
The relative frequencies of utilized and available velocities, depths and cover/substrate
types by sampling event are shown in Figures 9-29. Over 50% of available velocities among all
sampling events were less than 6 cm/s, and ranged from 0 to 65 cm/s. In general, the frequencies
of utilized velocities of all species and life stages showed similar trends to the frequencies of
environmental availability (Figures 9-15). However, the adult stages of all species utilized a
availability, principally among the juvenile stages of species. Juvenile black basses and spotted
51
Juvenile largemouth bass
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 9. Relative frequencies of water velocities available and utilized by juvenile largemouth
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
52
Adult largemouth bass
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 10. Relative frequencies of water velocities available and utilized by adult largemouth
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
53
Juvenile Suwannee bass
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 11. Relative frequencies of water velocities available and utilized by juvenile Suwannee
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
54
Adult Suwannee bass
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 12. Relative frequencies of water velocities available and utilized by adult Suwannee
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
55
Juvenile spotted suckers
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 13. Relative frequencies of water velocities available and utilized by juvenile spotted
suckers by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
56
Adult spotted suckers
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 14. Relative frequencies of water velocities available and utilized by adult spotted
suckers by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
57
American eels
Winter 2006 Winter 2007
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Relative frequency
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 15. Relative frequencies of water velocities available and utilized by American eels by
sampling event. Available velocities are represented by broken lines, and utilized velocities are
represented by solid lines.
58
suckers generally utilized shallower depths than adult stages (Figures 16, 18, 20), whereas adult
stages including American eels utilized deeper habitats as well as a greater range of depths
(Figures 17, 19, 21, 22). Available cover types among all sampling events were generally
dominated by boulder and rock outcroppings, snags, tree roots/cypress knees and organic debris
(Figures 23-29), (Refer to Table 2 for cover codes). Floating vegetation and algal mats were a
relatively abundant habitat in the summer and fall of 2007. Fish use of cover varied among
sampling events, but in general there were also disproportional uses of cover relative to available
cover types. American eels utilized boulder and limerock outcropp ings in greater frequency than
other cover types across all sampling events (Figure 29). Juvenile and adult stages of black
basses and spotted suckers always utilized snag and boulder habitats, however, juvenile fishes
utilized sand habitats with no cover and organic debris more frequently than adults, whereas
As a result of the missing 2005 year classes of black basses and spotted suckers, and
small sample sizes among several seasonal sampling events, the construction of habitat
suitability curves (HSC) within seasons to serve as statistical replicates was not possible.
However, since general similarities in relative use of the environmental variables within seasons
were present, data were pooled within seasons to derive one seasonal suitability curve per
variable, and per species life stage. In general, water velocities less than 30 cm/s were suitable
for all juvenile black basses and spotted suckers across all seasons, but typically selected
59
Juvenile largemouth bass
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 16. Relative frequencies of water depths available and utilized by juvenile largemouth
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
60
Adult largemouth bass
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 17. Relative frequencies of water depths available and utilized by adult largemouth bass
by sampling event. Available velocities are represented by broken lines, and utilized velocities
are represented by solid lines.
61
Juvenile Suwannee bass
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 18. Relative frequencies of water depths available and utilized by juvenile Suwannee
bass by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
62
Adult Suwannee bass
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 19. Relative frequencies of water depths available and utilized by adult Suwannee bass
by sampling event. Available velocities are represented by broken lines, and utilized velocities
are represented by solid lines.
63
Juvenile spotted suckers
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 20. Relative frequencies of water depths available and utilized by juvenile spotted
suckers by sampling event. Available velocities are represented by broken lines, and utilized
velocities are represented by solid lines.
64
Adult spotted suckers
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 21. Relative frequencies of water depths available and utilized by adult spotted suckers
by sampling event. Available velocities are represented by broken lines, and utilized velocities
are represented by solid lines.
65
American eels
Winter 2006 Winter 2007
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
Relative frequency
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
0.60 0.60
0.40 0.40
0.20 0.20
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 22. Relative frequencies of water depths available and utilized by American eels by
sampling event. Available velocities are represented by broken lines, and utilized velocities are
represented by solid lines.
66
Juvenile largemouth bass
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 23. Relative frequencies of cover/substrates available and utilized by juvenile largemouth
bass by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
67
Adult largemouth bass
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 24. Relative frequencies of cover/substrates available and utilized by adult largemouth
bass by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
68
Juvenile Suwannee bass
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 25. Relative frequencies of cover/substrates available and utilized by juvenile Suwannee
bass by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
69
Adult Suwannee bass
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 26. Relative frequencies of cover/substrates available and utilized by adult Suwannee
bass by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
70
Juvenile spotted suckers
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 27. Relative frequencies of cover/substrates available and utilized by juvenile spotted
suckers by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
71
Adult spotted suckers
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 28. Relative frequencies of cover/substrates available and utilized by adult spotted
suckers by sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
72
American eels
Winter 2006 Winter 2007
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00
Relative frequency
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 29. Relative frequencies of cover/substrates available and utilized by American eels by
sampling event. Available cover/substrate is represented by broken bars. Utilized
cover/substrate is represented by solid bars. Codes are listed in Table 2.
73
water velocities less than 6 cm/s (Figuress 30-32). Water velocities less than 30 cm/s were also
suitable for adult black basses and spotted suckers, but they tended to select water velocities
from 12 to 24 cm/s. Both juvenile and adult Suwannee bass selected relatively greater water
velocities than juvenile and adult largemouth bass, respectively, within the same seasons. The
water velocity suitability for American eels was also generally less than 30 cm/s, however, the
Juvenile fishes showed similar trends for depth, selecting shallower habitats across
seasons (Figures 34-36). Depths less than 240 cm were suitable for juvenile black basses and
spotted suckers, but they tended to select depths from 30 to 90 cm. By contrast, adult black
basses, adult spotted suckers and American eels selected a wider range of depths within and
among seasons, with overall suitability ranging from 60 to 450 cm (Figures 34-37). Within
black basses, juvenile largemouth bass selected greater depths in all seasons than juvenile
Suwannee bass, whereas no discernable trends were observed among the adults.
The selection of cover types among juvenile fishes was different. In the winter, juvenile
largemouth bass selected emergent grasses, juvenile Suwannee bass selected organic debris, and
juvenile spotted suckers selected overhanging vegetation (Figures 38-40), (Refer to Table 2 for
cover codes). In the spring, juvenile largemouth bass mostly selected falldowns, juvenile
Suwannee bass selected cobble/gravel habitats, and juvenile spotted suckers selected sandy flats
with no cover. In the summer, juvenile black basses selected similar habitats, primarily using
organic debris, floating vegetation/algae, and snag habitats, while spotted suckers continued to
select sandy substrates with no cover. In the fall, juvenile black basses again selected floating
74
Juvenile largemouth bass Adult largemouth bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Suitability Ratio
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 30. Water velocity suitability curves by season for juvenile and adult largemouth bass.
75
Juvenile Suwannee bass Adult Suwannee bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Suitability Ratio
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 31. Water velocity suitability curves by season for juvenile and adult Suwannee bass.
76
Juvenile spotted suckers Adult spotted suckers
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 32. Water velocity suitability curves by season for juvenile and adult spotted suckers. No
juveniles were collected in the fall seasons.
77
American eels
Winter
1.00
0.75
0.50
0.25
0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84
Spring
1.00
0.75
0.50
0.25
0.00
Suitability Ratio
6 12 18 24 30 36 42 48 54 60 66 72 78 84
Summer
1.00
0.75
0.50
0.25
0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84
Fall
1.00
0.75
0.50
0.25
0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 33. Water velocity suitability curves by season for American eels.
78
Juvenile largemouth bass Adult largemouth bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 34. Depth suitability curves by season for juvenile and adult largemouth bass.
79
Juvenile Suwannee bass Adult Suwannee bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 35. Depth suitability curves by season for juvenile and adult Suwannee bass.
80
Juvenile spotted suckers Adult spotted suckers
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Suitability Ratio
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 36. Depth suitability curves by season for juvenile and adult spotted suckers. No
juveniles were collected in the fall seasons.
81
American eels
Winter
1.00
0.75
0.50
0.25
0.00
30 90 150 210 270 330 390 450 510
Spring
1.00
0.75
0.50
0.25
0.00
Suitability Ratio
Summer
1.00
0.75
0.50
0.25
0.00
30 90 150 210 270 330 390 450 510
Fall
1.00
0.75
0.50
0.25
0.00
30 90 150 210 270 330 390 450 510
Depth (cm)
82
Juvenile largemouth bass Adult largemouth bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Substrate/cover code
Figure 38. Cover/substrate suitability plots by season for juvenile and adult largemouth bass.
Codes are listed in Table 2.
83
Juvenile Suwannee bass Adult Suwannee bass
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Substrate/cover code
Figure 39. Cover/substrate suitability plots by season for juvenile and adult Suwannee bass.
Codes are listed in Table 2.
84
Juvenile spotted suckers Adult spotted suckers
Winter Winter
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Spring Spring
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Summer Summer
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Fall Fall
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
Substrate/cover code
Figure 40. Cover/substrate suitability plots by season fo r juvenile and adult spotted suckers.
Codes are listed in Table 2.
85
American eels
Winter
1.00
0.75
0.50
0.25
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14
Spring
1.00
0.75
0.50
0.25
0.00
Suitability Ratio
1 2 3 4 5 6 7 8 9 10 11 12 13 14
Summer
1.00
0.75
0.50
0.25
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14
Fall
1.00
0.75
0.50
0.25
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14
Substrate/cover code
Figure 41. Substrate/cover suitability plots by season for American eels. Codes are listed in
Table 2.
86
debris, but Suwannee bass also selected undercut banks. Although not highly seasonally
selected, snags and roots/knees were always seasonally suitable and therefore similarly important
habitats.
Among adult black basses and spotted suckers, falldown and snag habitats were highly
selected in most cases, and utilized among all seasons with the exception of the summer by
spotted suckers (Figures 38-40). When available in the spring and fall, floating vegetation/algae
was also a suitable habitat with a high selection value. American eels highly selected
boulder/rock outcroppings in the spring and summer, bedrock with no cover in winter and
undercut banks in the fall (Figure 41). Falldowns were also highly selected by American eels in
the spring.
Overall, the seasonal suitability trends for velocity and depth by juvenile and adult black
basses and spotted suckers appeared similar. However, the seasonal cover/substrate suitabilities
were more variable, as were the trends for velocity, depth and cover by American eels. Despite
this fact, the seasonal suitabilities were combined to derive one suitability curve per variable per
species life stage for habitat simulation purposes. To accomplish this task, the ratios between
the relative frequencies of utilization and availability were summed across the seasonal variable
intervals, and divided by the maximum ratio to standardize the data from a scale of zero to one.
This approach was sensible considering the utilized and available sample sizes among seasons
for each species life stage were relatively similar. Therefore, the pooled dataset was not biased
towards a particular season. Additionally, the simulation of usable area of each species by
habitat suitability curves were the final criteria that were used in the PHABSIM analysis.
87
The pooled HSCs demonstrated that largemouth bass juveniles selected water velocities
less than 6 cm/s, while water velocities between 6 to 18 cm/s were moderately selected (Figure
42). Water velocities greater than 24 cm/s were not selected. Adult largemouth bass selected
water velocities from 0 to 24 cm/s, and selection decreased beyond 36 cm/s (Figure 42). Depth
selection by juvenile largemouth bass was limited to less than 120 cm, with an optimum
suitability depth of 60 cm (Figure 43). Depths greater than 120 cm were relatively not selected.
Adult largemouth bass utilized the range of available measured depths, and ranged from 30 to
500 cm, but depths in the range of 120 to 240 cm were mostly selected (Figure 43). The three
most selected cover/substrate types by juvenile largemouth bass were organic debris, floating
vegetation/algae and snag habitats (Figure 44). Adult largemouth bass typically selected
Suwannee bass juveniles most selected water velocities less than 6 cm/s, where suitability
values declined to zero beyond 24 cm/s (Figure 42). Adult Suwannee bass increasingly selected
velocities from 0 to 24 cm/s, where suitability declined beyond velocities above 36 cm/s. Depth
selection by juvenile Suwannee bass was highest below 90 cm, and selection decreased beyond
120 cm, while adults selected a wide range of depths peaking at 180 cm and then decreased to
420 cm (Figure 43). Juvenile Suwannee bass selected organic debris, roots/knees, undercut
banks, snags and floating vegetation/algae habitats, whereas adults selected falldowns,
As with juvenile black basses, juvenile spotted suckers most selected water velocities less
than 6 cm/s, where the suitability declined to zero beyond 36 cm/s (Figure 42). Adult spotted
suckers increasingly selected velocities from 0 to 30 cm/s, and velocity suitability declined
abruptly beyond 30 cm/s. Juvenile spotted suckers most selected depths between 60 and 120 cm,
88
Largemouth bass juveniles Largemouth bass adults
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84 6 12 18 24 30 36 42 48 54 60 66 72 78 84
American eels
1.00
0.75
0.50
0.25
0.00
6 12 18 24 30 36 42 48 54 60 66 72 78 84
Velocity (cm/s)
Figure 42. Pooled velocity (cm/s) habitat suitability curves across all collection dates and
seasons on the Withlacoochee River, FL.
89
Largemouth bass juveniles Largemouth bass adults
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
30 90 150 210 270 330 390 450 510 30 90 150 210 270 330 390 450 510
American eels
1.00
0.75
0.50
0.25
0.00
30 90 150 210 270 330 390 450 510
Depth (cm)
Figure 43. Pooled depth (cm) habitat suitability curves across all collection dates and seasons on
the Withlacoochee River, FL.
90
Largemouth bass juveniles Largemouth bass adults
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
Suitability Ratio
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
1.00 1.00
0.75 0.75
0.50 0.50
0.25 0.25
0.00 0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14 1 2 3 4 5 6 7 8 9 10 11 12 13 14
American eels
1.00
0.75
0.50
0.25
0.00
1 2 3 4 5 6 7 8 9 10 11 12 13 14
Cover/substrate code
Figure 44. Pooled cover/substrate habitat suitability plots across all collection dates and seasons
on the Withlacoochee River, FL.
91
whereas adult spotted suckers used a wide range of depths under 390 cm, with a suitability peak
between 180 to 240 cm (Figure 43). Juvenile spotted suckers selected over- hanging vegetation
and sand cover/substrate types whereas adults selected falldown, snag and sand cover/substrate
American eels selected water velocities from 0 to 30 cm/s, and did not select velocities
greater than 36 cm/s (Figure 42). Eels utilized a wide range in depths (Figure 43), and
predominantly selected undercut banks, boulder/rock outcroppings, and root/knee cover types
(Figure 44).
The output of the PHABSIM model provided estimates of the amount of usable area at
each of the habitat reaches for the targeted species over a range of discharges, based on the
habitat suitability criteria and the habitat reach transects. WUA estimates at each discharge were
summed across each habitat reach to provide a composite WUA estimate that was representative
of shoal and run mesohabitats within the study area for each life stage of the targeted species.
All WUA estimates for each habitat reach are available by life stage and species in Appendix 6.
In general, the physical area available for adult black basses and spotted suckers were
greater than their juvenile life stages at a given discharge at each of the habitat transects sampled.
The range of usable area for juvenile largemouth bass was 9,700 ft2 /1000 linear ft to 47,000
ft2 /1000 linear ft across the simulated discharges (Figure 45). Maximum usable area for juvenile
largemouth bass was available at low discharges (135 cfs), and then steadily declined as
discharge increased. Conversely, the WUA for adult largemouth bass increased from low to
intermediate discharges reaching a maximum of 62,600 ft2 /1000 linear ft at 475 cfs, followed by
92
Largemouth bass
70000
WUA (sq ft / 1000 linear ft)
60000
50000
40000 Adults
30000 Juveniles
20000
10000
0
135
160
200
250
300
350
400
525
700
950
1200
Discharge (cfs)
Figure 45. Composite weighted usable area per incremental discharge available to juvenile and
adult stages of largemouth bass in the study area of the Withlacoochee River, FL.
93
declining usable areas as discharges increased (Figure 45). Estimates of usable area for adult
largemouth bass ranged from 35,700 ft2 /1000 linear ft to 62,600 ft2 /1000 linear ft.
Juvenile Suwannee bass displayed similar trends to juvenile largemouth bass in usable
area, with the model predicting higher values of usable area at low discharges and declining as
discharges increased. However, the slope for juvenile Suwannee bass was less precipitous than
the curve for juvenile largemouth bass, and the predicted usable area for juvenile Suwannee bass
was greater across all discharge values in comparison to juvenile largemouth bass. Usable area
for juvenile Suwannee bass ranged from 23,100 ft2 /1000 linear ft to 56,000 ft2 /1000 linear ft and
was more than double at high discharge values compared to juvenile largemouth bass (Figure
46).
Adult Suwannee bass likewise showed trends in usable area that were similar to adult
largemouth bass. The usable area increased from low to intermediate discharges, and then
decreased at high discharges (Figure 46). However, the WUA estimates were greater than
estimates predicted for largemouth bass across all discharges. Additionally, the discharge in
which the asymptote was observed was greater for adult Suwannee bass, providing 127,500
The trend in usable area for juvenile spotted suckers was again similar to those observed
in juvenile black basses. Usable area was generally maximized at low discharges and declined
steadily as discharge increased (Figure 47). Usable area predictions ranged from 6,100 ft 2 /1000
linear ft to 39,000 ft2 /1000 linear ft, which were the lowest estimates among the target species
Usable area predictions for adult spotted suckers were the highest among the target
species and life stages. Usable area values ranged from 100,000 ft2 /1000 linear ft to 202,000
94
Suwannee bass
140000
WUA (sq ft / 1000 linear ft)
120000
100000
80000 Adults
60000 Juveniles
40000
20000
0
135
160
200
250
300
350
400
525
700
950
1200
Discharge (cfs)
Figure 46. Composite weighted usable area per incremental discharge available to juvenile and
adult stages of Suwannee bass in the study area of the Withlacoochee River, FL.
95
Spotted suckers
250000
WUA (sq ft / 1000 linear ft)
200000
150000
Adults
Juveniles
100000
50000
0
135
160
200
250
300
350
400
525
700
950
1200
Discharge (cfs)
Figure 47. Composite weighted usable area per incremental discharge available to juvenile and
adult stages of spotted suckers in the study area of the Withlacoochee River, FL.
96
ft2 /1000 linear ft (Figure 47). The usable area was predicted to be lowest at low discharges and
increasing to maximum usable area at 700 cfs, followed by declines in usable area when
The usable area predictions for American eels were highly variable across habitat reaches
and no discernable trends were observed. However, the summed estimates pr edicted the WUA
to trend similar to the adult stages of the other target species. Estimates were greatest and
increased from low discharges to a maximum of 99,300 ft2 /1000 linear ft at 340 cfs, and then
The graphical results of the time-series analyses across all habitat reaches for each
species and life stage are in Appendix 7. As with the WUA estimates, calculated values of
habitat gains/losses pertaining to percent reductions in discharge were summed across habitat
reaches to derive a composite assessment of habitat gains or losses in the study area representing
shoal and run mesohabitats. Largemouth bass juveniles are predicted to gain habitat across all
months in both time periods as discharge is reduced 10% to 40% from existing conditions
(Figure 49). Largemouth bass adults also show habitat gains across all months with increasing
percent reductions of discharge (Figure 50). Greatest habitat gains are observed in March and
April, whereas relatively lower gains are predicted in October through December.
Suwannee bass juveniles showed habitat gains across all months with reductions in
discharge from 10 to 40% (Figure 51). In comparison to juvenile largemouth bass, Suwannee
bass are expected to receive relatively less habitat gains as discharge is reduced. Adults similarly
showed habitat gains across nearly all percent reductions of discharge (Figure 52). Habitat loss
(-2%) was only observed at 40% reduction in discharge in November during the
97
American eels
120000
WUA (sq ft / 1000 linear ft)
100000
80000
60000 Adults
40000
20000
0
135
160
200
250
300
350
400
525
700
950
1200
Discharge (cfs)
Figure 48. Composite weighted usable area per incremental discharge available to adult stages
of American eels in the study area of the Withlacoochee River, FL.
98
Largemouth bass juveniles
1940-1969
80%
Habitat gains / losses
70%
60%
50% 10% reduction
20% reduction
40%
30% reduction
30% 40% reduction
20%
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
80%
Habitat gains / losses
70%
60%
50% 10% reduction
20% reduction
40%
30% reduction
30% 40% reduction
20%
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 49. Composite time-series analysis for juvenile largemouth bass of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
99
Largemouth bass adults
1940-1969
60%
Habitat gains / losses
50%
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
60%
Habitat gains / losses
50%
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 50. Composite time-series analysis for adult largemouth bass of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
100
Suwannee bass juveniles
1940-1969
50%
Habitat gains / losses
40%
10% reduction
30%
20% reduction
30% reduction
20%
40% reduction
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
50%
Habitat gains / losses
40%
10% reduction
30%
20% reduction
30% reduction
20%
40% reduction
10%
0%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 51. Composite time-series analysis for juvenile Suwannee bass of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
101
Suwannee bass adults
1940-1969
45%
40%
Habitat gains / losses
35%
30% 10% reduction
25%
20% reduction
20%
30% reduction
15%
10% 40% reduction
5%
0%
-5%
November
June
January
August
March
July
February
September
December
April
May
October
45%
40%
Habitat gains / losses
35%
30% 10% reduction
25%
20% reduction
20%
30% reduction
15%
10% 40% reduction
5%
0%
-5%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 52. Composite time-series analysis for adult Suwannee bass of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
102
Spotted suckers juveniles
1940-1969
140%
Habitat gains / losses
120%
100%
10% reduction
80% 20% reduction
60% 30% reduction
40% reduction
40%
20%
0%
November
June
January
August
March
July
February
September
December
April
May
October
140%
Habitat gains / losses
120%
100%
10% reduction
80% 20% reduction
60% 30% reduction
40% reduction
40%
20%
0%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 53. Composite time-series analysis for juvenile spotted suckers of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
103
Spotted suckers adults
1940-1969
60%
Habitat gains / losses
50%
40%
10% reduction
30% 20% reduction
20% 30% reduction
40% reduction
10%
0%
-10%
November
June
January
August
March
July
February
September
December
April
May
October
60%
Habitat gains / losses
50%
40%
10% reduction
30% 20% reduction
20% 30% reduction
40% reduction
10%
0%
-10%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 54. Composite time-series analysis for adult spotted suckers of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
104
American eel adults
1940-1969
45%
40%
Habitat gains / losses
35%
30% 10% reduction
25% 20% reduction
20% 30% reduction
15% 40% reduction
10%
5%
0%
November
June
January
August
March
July
February
September
December
April
May
October
45%
40%
Habitat gains / losses
35%
30% 10% reduction
25% 20% reduction
20% 30% reduction
15% 40% reduction
10%
5%
0%
November
June
January
August
March
July
February
September
December
April
May
October
Figure 55. Composite time-series analysis for adult American eels of habitat gains/losses
associated with 10-40% reductions in monthly discharge.
105
1940-1969 period, and in October (1%) and November (1%) during the 1970-1999 period.
Greatest habitat gain was observed in March and April, while least habitat change was observed
Similar to juvenile stages of black basses, juvenile spotted suckers showed habitat gains
across all levels of discharge reductions during both time periods (Figure 53). Greatest habitat
gains are predicted during March and April, while lesser gains are shown during October and
November. Adult stages showed a peak in habitat gains across all percent reductions during
March and April with relatively minor gains in other months (Figure 54). Habitat loss was
predicted only above 20% reduction in discharges during both time periods during the fall
months, with the maximum habitat loss (8%) occurring at during the 1940-1969 simulation.
The results for American eels were similar to those of the other adult stages. Habitat
gains were predicted over all percentages of discharge reductions, with maximum gains
occurring during March and April and least habitat change occurring during the fall months
(Figure 55).
Discussion
It is important that biological tools become an integral component of the process for
establishing minimum flows and levels in Florida streams. Most methods currently utilized for
provide for the protection of aquatic biota and ecological processes, and ensure ample water
supply for consumptive users. While current methods give the appearance of being adequate in
their applications to-date, it is logical to assume that ecological tools would be most suitable for
ensuring the protection of ecological integrity. Ecological approaches are needed to fine-tune
106
current MFL methodology to the unique biotic attributes of individual stream systems. As the
analyses presented herein demonstrate, each individual stream system can have unique
ecological indicator species and functional guilds that, in turn, have specific streamflow and
water level needs. Setting MFLs based solely upon hydrological and geological assumptions can
result in unmet ecological requirements of species critical to stream ecosystem function and
integrity.
Results of our analyses show that fine particle size organic matter (FPOM) and attached
algae drive the food chain in the Withlacoochee River study area. The gathering, filter ing, and
scraping invertebrate taxa that depend upon these food resources accounted for from 68 to 85
percent of the total invertebrate organisms collected from the six sampled habitat types. Many of
these gathering, filtering, and scraping taxa are important food web components and/or are
integral to ecosystem metabolism processes. In general, these taxa are most abundant in the
snag, cobble/gravel riffle, and tree root system habitats and they require flow to supply their food
(e.g. net-spinning caddisflies) and/or to oxygenate their gills (e.g. baetid mayflies). Many of
these taxa also have long life spans (1 year or longer) and, hence, require that their habitats be
inundated for long time periods. The snags and root systems where many of these taxa reach
their greatest abundances are located above the stream bottom, and, consequently, are among the
first habitats to be dewatered under receding water level conditions. Given the ecological
importance of the snag and tree root-dwelling fishes and invertebrates, it is critical that above-
The cobble/gravel riffle habitat is also ecologically critical because of its elevation above
other bottom habitats and because of its structural complexity (and consequent habitat
complexity). Cobble/gravel riffles in the Withlacoochee are located upon the bedrock shoals that
107
serve as hydraulic streamflow controls. Flow reductions and dewatering of this habitat for
prolonged periods of time will result in reduced available habitat area and profound structural
The flow (current velocity) needs of many invertebrate taxa are quite specific, as
demonstrated by the current velocity habitat suitability curves and weighted usable habitat areas
presented in Appendices 3 and 4. The Withlacoochee River invertebrate taxa found to be most
responsive to flows were the nonbiting midges Parakiefferiella sp. and Pseudochironomus sp.
Other taxa that were substantially affected by flow included: total Chironomidae, the individual
chironomid species Cricotopus bicinctus and Tvetenia vitracies, total Baetidae (Ephemeroptera),
Corydalus cornutus. We believe this suite of nine invertebrate taxa to be the most suitable
biological indicator of flow response in the Withlacoochee system. Monitoring of this suite in
the water- level sensitive snag, cobble/gravel riffle, and tree root system habitats will provide a
valuable tool for the evaluation of flow and water level reduction impacts. The community
quality descriptors total taxa, Shannon diversity, and total organisms did not exhibit strong flow
responses, and, along with Hydracarina, Stenelmis sp., Stempellina sp. A, and Tanytarsus sp. C,
We believe that the PHABSIM simulation model and time series analysis of PHABSIM
results provide a useful tool for evaluating the potential ecological consequences of flow and
water level reductions in many Florida stream systems. Our invertebrate community PHABSIM
results indicate that maintaining adequate flow throughout the autumn months of September
taxa in the Withlacoochee River study area (Tables 11 - 13, Appendix 5). We demonstrated that
108
flow reductions of 30 to 40 percent from the monthly mean during the autumn months would
adversely affect populations of dominant invertebrate indicator taxa and would probably result in
long-term negative impacts to invertebrate community structure within the study area. To
maintain optimal habitat area and population levels of the ecologically important net-spinning
caddisfly taxa (e.g. total Hydropsychidae and Hydropsyche sp.), minimum discharges of 340 cfs
are required. To maintain optimal habitat area and population levels of the more flow sensitive
chironomid taxa (e.g. Pseudochironomus sp. and Parakiefferiella sp.), discharges exceeding
Conversely, the fish species selected for this study were not overly sensitive to stream
flow reductions based on results of the WUA and time-series analyses simulations. In fact,
reductions in stream flow mostly led to habitat gains in both juvenile and adult stages of the
target species examined based on the time-series analysis. However, several trends emerged
from the results that were life-stage specific rather than species-specific. Juvenile black basses
and spotted suckers all selected slow water velocities and shallow depths. Conversely, water
velocity and depth selection among adults tended to be more normally distributed, reaching
maximum suitabilities at intermediate velocities and depths, while extreme ends of the habitat
variables were relatively less suitable. This trend was logical since the drag associated with
current velocity is known to influence fish endurance, foraging success (Schae fer et al. 1999) and
swimming performance (Deegan et al. 2005), where current velocity occupancy is generally
related to physical body size. Size-specific habitat use has also been commonly documented in
many stream fishes due to reasons associated with competition (Freeman and Stouder 1989) and
predation avoidance (Power 1987, Schlosser 1987). Consequently, the amount of usable area
available to juveniles in the study area of the Withlacoochee River decreased as discharge
109
increased, whereas adults generally obtained optimum usable area between discharges of 500-
700 cfs, followed by decreasing area as discharge increased. Examination of the mean monthly
discharge at the Lee gage indicated that most flows were typically greater than 1,100 cfs.
Therefore existing conditions more reflect the trends and values of the WUA estimates at the
Despite commonalities among species and life stages, differences within species were
also noted. Juvenile and adult Suwannee bass selected relatively greater water velocities than
juvenile and adult largemouth bass, respectively. Conversely, juvenile largemouth bass selected
greater depths than juvenile Suwannee bass. Cover/substrate types were commonly selected
among adult basses, but boulder/rock outcropping habitats were more suitable to Suwannee bass
than largemouth bass. Thus, WUA estimates were higher for Suwannee bass than for
largemouth bass across all discharges, and suggest that Suwannee bass may indeed possess a
greater affinity to flow. Although, the time-series analysis revealed that black basses would
generally not be negatively impacted by reductions in stream flow based on the composite
calculations, comparisons among juveniles suggests less habitat gains by Suwannee bass than by
largemouth bass, and greater potential habitat losses by Suwannee bass adults at 40% reduction
in stream flows than for largemouth bass adults. Exaggeration of this simulation could
potentially result in the conversion of habitats more suitable for largemouth bass than for
Suwannee bass with increased flow reductions, causing an ecological shift in the community
composition.
Based on the collective examinations of the WUA and time-series for each fish species
and life stage, flow recommendations are difficult. The directive from the Florida Statue to the
five water management districts is to set levels below which significant harm may occur.
110
Several Districts have used 15% as a benchmark above which habitat loss is considered
significant, and below which habitat loss is considered acceptable. Under all composite time-
series presented, no more than 10% habitat is lost even under a 40% reduction in flow from
existing conditions. Thus, based on these criteria for significant harm, this reduction in stream
flow may be acceptable to the fish species selected. A more conservative approach may be to
examine each of the habitat reaches individually and inspect the habitat and passage losses that
are predicted to occur with flow reductions. Under this scenario, Reach 1 (referred to shoal 1 in
Appendix 7) exhibited the greatest loss of habitat to Suwannee bass adults, predicting losses in 6
months out of the year under 40% flow reductions, with a maximum of 26% loss in November
during the 1940-1969 time period, and a maximum 21% loss in October and November during
the 1970-1999 time period. Habitat losses were also expressed in these months from 10% to
30% reductions in stream flow, albeit to a lesser degree, and not exceeding 15% habitat loss.
Adult stages of largemouth bass and spotted suckers similarly exhibit habitat loss in Reach 1 that
exceeded 15% during the fall months. Because Reach 1 is the most upstream habitat transect
likely provide the most protective flows and levels management to the target fish species in all
downstream habitats of the Withlacoochee River. However, those fishes not encompassed by the
habitat suitabilities of the target species described within may not be fully protected under this
scenario, and care should be taken to ensure passage of fishes such as the anadromous gulf
The fact that these fish species were not overly sensitive to stream flow reductions may
be in part a function of selecting run and pool-dwelling species for PHABSIM analysis, as
opposed to species that occupy shoal or riffle habitats. Since shoal and riffle habitats are the first
111
to be de-watered with reduced flows, fish species that typically occupy these habitats would
likely show the greatest losses in usable habitat. Peninsular Floridas fish fauna generally lack
obligate riverine species, and are more composed of facultative riverine and lacustrine species
that utilize more general and ubiquitous habitats. Another function that may have led to a bias
towards selection of slow velocities and shallow depths was the fish sampling approach. Over
50% of the available velocities among all sampling events were less than 6 cm/s. Although fish
collections were made seasonally, specific effort was made to collect fishes during low flow
fishes over a range of discharges is advised. Because the intention of this study was for MFL
purposes, we emphasized the habitat selection during low flow conditions, and felt sampling the
target fish species during median and high discharge events was both unnecessary and unfeasible
Conclusions
Results of our community- level and PHABSIM analyses suggest that streamflow
reductions beyond the range of natural seasonal variation could significantly impact the
ecological integrity of the Withlacoochee system. Principal conclusions from this research were:
and scraping taxa that are dependent upon fine particle sized detritus and attached
cobble/gravel riffle habitat and in the above-bottom snag and tree root system habitats.
Snag and tree root systems were also important habitats selected by the target fish
species. Monitoring and protection of these habitats is critical since they harbor sessile,
112
long- lived, ecologically important communities and are the first habitats to be de-watered
2. Optimum habitat areas for most dominant and/or flow sensitive invertebrate taxa are
available at discharges ranging from 340 to 1300 cfs. 5. Autumn months are critical for
invertebrate taxa and the most vulnerable months for fish species.
3. Flow reductions of 30 to 40 percent from the monthly mean during the autumn months of
4. The targeted fish species for PHABSIM modeling do not appear to be suitable candidates
mostly habitat gains for the selected fish species, and a 40% red uction in flow was
necessary to produce about a 10% habitat loss based on composite reach calculations.
5. Flow reductions could provide conditions more suitable for largemouth bass than
Suwannee bass, and invertebrate composition change due to flow reduction would likely
113
LITERATURE CITED
ALLAN, J.D. 1995. Stream ecology: structure and function of running waters. Kluwer
Academic Publishers. Dordrecht, the Netherlands. 388 pp.
ANNEAR T., I. CHISHOLM, H. BEECHER, A. LOCKE, and 12 other authors. 2004. Instream
flows for riverine resource stewardship, revised edition. Instream Flow Council,
Cheyenne, WY. 268 pp.
BAIN, M.B., J.T. FINN, and H.E. BOOKE. 1988. Streamflow regulation and fish community
structure. Ecology 69:382-392.
BARBOUR, M.T., J. GERRITSEN, B.D. SNYDER, and J.B. STRIBLING. 1999. Rapid
bioassessment protocols for use in streams and wadeable rivers: periphyton, benthic
macroinvertebrates, and fish, second edition. EPA 841-B-99-002. U.S. Environmental
Protection Agency; Office of Water; Washington, D.C.
BERNDT, M.P., E.T. OAKSFORD, G.L. MAHON, and W. SCHMIDT. 1998. Groundwater. In
E.A. Fernald and E.D. Purdum (eds.). Water Resources Atlas of Florida. Institute of
Science and Public Affairs, Florida State University. 312 pp.
BERNER, L., AND M.L. PESCADOR. 1988. The mayflies of Florida, revised edition.
University Presses of Florida, Tallahassee and Gainesville. 415pp.
BOVEE, K.D. 1986. Development and evaluation of habitat suitability criteria for use in the
instream flow incremental methodology. Instream Flow Information Paper 21. U.S. Fish
and Wildlife Service Biological Report 86(7). 235 pp.
BOVEE, K.D. 1997. Data collection procedures for the physical habitat simulation system.
U.S. Geological Survey, BRD, Mid-Continent Science Center, Ft. Collins, CO. 146 pp.
BOVEE, K.D., J.A. GORE, AND A.J. SILVERMAN. 1978. Field-testing and adaptation of a
methodology to measure instream values in the Tongue River, Northern Great Plains
(NGP) Region. U.S. Environmental Protection Agency, EPA-908/4-78-004A. 465 pp.
BOVEE, K.D., B.L. LAMB, J.M. BARTHOLOW, C.B. STALNAKER, J. TAYLOR, and J.
HENRIKSEN. 1998. Stream habitat analysis using the instream flow incremental
methodology. U.S. Geological Survey, BRD Information and Technology Report
USGS/BRD-1998-0004. viii + 131 pp.
114
CALOW, P., and G.E. PETTS. 1992. The rivers handbook: hydrological and ecological
principals. Vol. 1. Blackwell Scientific Publications, Boston, MA. 526 pp.
CALOW, P., and G.E. PETTS. 1994. The rivers handbook: hydrological and ecological
principals. Vol. 2. Blackwell Scientific Publications, Boston, MA. 523 pp.
CUMMINS, K.W. 1973. Trophic relations of aquatic insects. Annual Review of Entomology
18:183-206.
CUMMINS, K.W. 1974. Structure and function of stream ecosystems. Bioscience 24:631-641.
CUMMINS, K.W., R.W. MERRITT, and M.B. BERG. 2008. Ecology and distribution of
aquatic insects. In R.W. Merritt, K.W. Cummins, and M.B. Berg (eds.). An introduction
to the aquatic insects of North America, fourth edition. Kendall/Hunt Publishing Co.,
Dubuque, IA. xvi + 39 color plates + 1158 pp.
EDINGTON, J.M. 1968. Habitat preferences in net-spinning caddis larvae with special
reference to the influence of water velocity. Journal of Animal Ecology 37:675-692.
FREEMAN, M.C. and D.J. STOUDER. 1989. Intraspecific interactions influence size specific
depth distribution in Cottus bairdi. Environmental Biology of Fishes 24:231-236.
GORE, J.A. 1987. Development and applications of macroinvertebrate instream flow models
for regulated flow management. In: J.F. Craig and J.B. Kemper, eds. Regulated
streams: advances in ecology. Plenum Press, New York. pp. 99-115.
GORE, J.A. and R.D. JUDY. 1981. Predictive models of benthic macroinvertebrate density for
use in instream flow studies and regulated flow management. Can. Journ. Fish. Aquat.
Sci. 38:1363-1370.
GORE, J.A., J.B. LAYZER, and J. MEAD. 2001. Macroinvertebrate instream flow studies after
20 years: a role in stream management and restoration. Regulated Rivers: Research and
Management 17:527-542
KARR J. R. 1981. Assessment of biotic integrity using fish communities. Fisheries. 6(6):2127.
KINGSOLVING, A.D., and M.B. BAIN. 1993. Fish assemblage recovery along a riverine
disturbance gradient. Ecological Applications 3:531-544.
115
KREBS, C.J. 1999. Ecological methodology, second edition. Addison Wesley Longman, New
York.
HAND, J., J. COL, and L. LORD. 1996. Northeast Florida District Water Quality Assessment
1996 305 (b) Technical Appendix. Bureau of Surface Water Management, Florida
Department of Environmental Protection.
HAWKINS, C.P. and J.R. SEDELL. 1981. Longitudinal and seasonal changes in functional
organization of macroinvertebrate communities in four Oregon streams. Ecology
62(2):387-397.
HYNES, H.B.N. 1970. The ecology of running waters. University of Toronto Press.
JACOBI, G.Z. 1978. An inexpensive circular sampler for collecting benthic macroinvertebrates
in streams. Archive fr Hydrobiologia 83:126-131.
MARELLA, R.L. 2008. Water use in Florida, 2005 and trends 1950-2005. U.S. Geological
Survey Fact Sheet 2008-3080. Available at:
http://fl.water.usgs.gov/infodata/wateruse.html
MERRITT, R.W., K.W. CUMMINS, M.B. BERG, J.A. NOVAK, M.J. HIGGINS, K.J.
WESSEL, and J.L. LESSARD. 2002. Development and application of a
macroinvertebrate functional- group approach in the bioassessment of remnant river
oxbows in southwest Florida. Journal North American Benthological Society 21(2):290-
310.
MERRITT, R.W., V.H. RESH, K.W. CUMMINS, and D.P. BATZER. 2008. Sampling aquatic
insects. In R.W. Merritt, K.W. Cummins, and M.B. Berg (eds.). An introduction to the
aquatic insects of North America, fourth edition. Kendall/Hunt Publishing Co., Dubuque,
IA. xvi + 39 color plates + 1158 pp.
MCCUNE, B. and M.J. MEFFORD. 1999. PC-ORD. Multivariate analysis of ecological data,
Version 5. MjM Software Design, Gleneden Beach, OR.
MILHOUSE, R.T., D.L. WEGNER, and T. WADDLE. 1984. Users guide to the physical
habitat simulation system. U.S. Fish and Wildlife Service, FWS/OBS 81/43.
MILHOUSE, R.T., M.A. UPDIKE, and D.M. SCHNEIDER. 1989. Physical habitat simulation
system reference manual, version II. U.S. Fish and Wildlife Service Biological Report
89.
MILLER D. L., and thirteen coauthors. 1988. Regional applications of an index of biotic
integrity for use in water resource management. Fisheries. 13(5):1220.
PIELOU, E.C. 1977. Ecological diversity. John Wiley and Sons, New York.
116
POFF, N.L. AND J.V. WARD. 1989. Implications of streamflow variability and predictability
for lotic community structure: a regional analysis of streamflow patterns. Canadian
Journal of Aquatic Sciences 46:1805-1818.
POFF, N.L., J.D. ALLAN, M.B. BAIN, J.R. KARR, K.L. PRESTEGAARD, B.D. RICHTER,
R.E. SPARKS, AND J.C. STROMBERG. 1997. The natural flow regime, a paradigm
for river conservation and restoration. Bioscience 47(11):769-784.
POWER, M.E. 1987. Predator avoidance by grazing fishes in temperate and tropical streams:
importance of stream depth and prey size, p 333-353. In W.C. Kerfoot and A. Sih (Eds.).
Predation: direct and indirect impacts on aquatic communities. University Press of New
England, Hanover, N.H.
POWERS, C.F. and A. ROBERTSON. 1967. Design and evaluation of an all-purpose benthos
sampler, pp. 126-131. In J.C. Ayers and D.C. Chandler (eds). Studies on the environment
and eutrophication of Lake Michigan. University of Michigan Great Lakes Research
Division Special Report 30.
RAO, D.V. and D.A. CLAPP. 1996. Preliminary evaluation of the impacts of spring discharge
reductions on the flows of receiving waters and natural systems, central Florida.
SJRWMD Special Publication SJ96-SP3. Palatka, FL. 71 pp.
RICHTER, B.D., J.V. BAUMGARTNER, J. POWELL, and D.P. BRAUN. 1996. A method for
assessing hydrologic alteration within ecosystems. Conservation Biology 10:1163-1174.
RICHTER, B.D, J.V. BAUMGARTNER, R. WIGINGTON, and D.P. BRAUN. 1997. How
much water does a river need? Freshwater Biology 37:231-249.
RESH, V.H., A.V. BROWN, A.P. COVICH, M.E. GURTZ, H.W. LI, G.W. MINSHALL, S.R.
REICE, A.L. SHELDON, J.B. WALLACE, and R. WISSMAR. 1988. The role of
disturbance in stream ecology. Journal of the North American Benthological Society
7(4)433-455.
SCHLOSSER, I.J. 1985. Flow regime, juvenile abundance, and the assemblage structure of
stream fishes. Ecology 66:1484-1490.
SCHLOSSER, I.J. 1987. The role of predation in age and size related habitat use by stream
fishes. Ecology 66: 651-659.
117
SCOTT, T.M, J.M. LLOYD, and G. MADDOX. 1991. Floridas ground water quality
monitoring program hydrogeological framework. Florida Geological Survey Special
Publication No. 32.
SIMPSON, K.W. and R.W. BODE. 1980. Common larvae of Chironomidae (Diptera) from
New York state streams and rivers. Bulletin No. 439. New York State Museum. 105 pp.
SMITH, D.I. AND P. STOPP. 1978. The river basin: an introduction to the study of hydrology.
Cambridge University Press, London. 120 pp.
SMITH, S.K. 2005. Florida population growth: past, present, and future. University of Florida
Dept. of Economic and Business Research.
STATZNER, B, J.A. GORE, and V.H. RESH. 1988. Hydraulic stream ecology: observed
patterns and potential applications. Journal of the North American Benthological Society
7(4)307-360.
VERGARA, B.A., ed. 1994. Water supply needs and sources assessment, 1994, St. Johns River
Water Management District. SJRWMD Technical Publication SJ94-7. Palatka, FL.
132pp.
WARD, J.V., and J.A. STANFORD. 1995. Ecological connectivity in alluvial river ecosystems
and its disruption by flow regulation. Regulated Rivers 11:105-119.
WARREN, G.L., T.J. FERRING, and D.A. HOHLT. 2006. Aquatic invertebrate communities
at five locations in the Alafia River watershed, Hillsborough County, Florida. Final
Report to Southwest Florida Water Management District. Florida Fish and Wildlife
Research Institute Library No. F6306-02-05-F.
WARREN, G.L., D.A. HOHLT, C.E. CICHRA, and D. VAN GENECHTEN. 2000. Fish and
aquatic invertebrate communities of the Wekiva and Little Wekiva Rivers: A baseline
evaluation in the context of Floridas minimum flows and levels statutes. Special
Publication SJ2000-SP4. St. Johns River Water Management District, Palatka, FL.
118
APPENDIX 1. Mean (no m-2), coefficient of variation (cv), and percent composition
(%) of aquatic invertebrate taxa collected from areally dominant habitats in the northern
Withlacoochee River, Florida, study area, 2005 - 2007.
119
APPENDIX 1. Mean (no m-2), coefficient of variation (cv), and percent composition (%) of aquatic invertebrate taxa collected from areally
dominant habitats in the northern Withlacoochee River, Florida, study area, 2005 - 2007.
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv)
Taxon % % % % % %
Cnidaria
Hydrozoa
Cordylophora lacustris P P P P P ---
Nemertea
Tetrastemmatidae
Prostoma sp. 99 (1.36) 214 (2.9) 642 (1.11) 27 (2.03) 107 (1.34) 19 (1.73)
0.3 1.6 1.2 0.1 0.2 0.1
Nematoda 140 (1.22) 284 (1.71) 317 (1.03) 512 (2.24) 372 (1.26) 261 (1.00)
0.4 2.1 0.6 1.7 0.6 1.5
Annelida
Aphanoneura
Aeolosomatidae
Aeolosoma sp. --- 2 (4.36) 5 (4.24) --- 17 (2.64) ---
<0.1 <0.1 <0.1
Oligochaeta (total) 5,003 (1.09) 2,802 (1.33) 4,854 (1.23) 582 (1.53) 3,317 (1.45) 319 (0.59)
12.9 20.5 8.8 2.0 5.5 1.8
120
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Naididae (total) 369 (2.07) 753 (3.87) 101 (1.82) 394 (2.16) 791 (0.91) 251 (0.79)
0.9 5.5 0.2 1.3 1.3 1.4
Dero digitata complex --- 215 (3.79) --- --- --- ---
1.6
121
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Nais behningi 48 (2.19) 10 (4.93) 16 (1.80) 188 (2.16) 184 (1.11) 24 (1.26)
0.1 0.1 <0.1 0.6 0.3 0.1
Nais communis complex 89 (1.93) 2 (4.36) 40 (2.26) 139 (2.60) 296 (1.69) 212 (0.85)
0.2 <0.1 0.1 0.5 0.5 1.2
122
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Slavina appendiculata <1 (3.61) --- --- <1 (5.48) 44 (1.44) ---
<0.1 <0.1 0.1
123
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
124
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
unidentifiable immature Oligochaeta 4 (3.61) --- 5 (3.20) <1 (5.48) --- ---
<0.1 <0.1 <0.1
Hirudinia
Glossiphoniidae <1 (3.61) --- 5 (3.76) --- --- 10 (1.73)
<0.1 <0.1 0.1
Mollusca
Gastropoda (total) 107 (1.62) 246 (2.08) 835 (1.36) 92 (1.75) 1,679 (1.09) 48 (0.35)
0.3 1.8 1.5 0.3 2.8 0.3
Micromenetus sp. <1 (3.61) --- --- <1 (5.48) 56 (3.87) ---
<0.1 <0.1 0.1
125
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Lymnaeidae
Pseudosuccinea columella 2 (3.61) --- 17 (3.90) --- --- ---
<0.1 <0.1
Pleuroceridae
Elimia floridensis 35 (1.74) 179 (2.33) 114 (1.73) 5 (3.08) 558 (2.51) ---
0.1 1.3 0.2 <0.1 0.9
Viviparidae
Viviparus georgianus --- --- 1 (4.24) 1 (5.48) 13 (2.74) ---
<0.1 <0.1 <0.1
126
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
unidentifiable imm. Gastropoda 48 (2.43) 25 (1.99) 260 (1.38) 36 (2.00) 100 (2.20) ---
0.1 0.2 0.5 0.1 0.2
Pelecypoda 131 (1.55) 2,045 (1.81) 665 (1.14) 45 (2.00) 342 (2.57) 155 (0.78)
0.3 15 1.2 0.2 0.6 0.9
Corbiculidae
Corbicula fluminea 22 (1.76) 779 (2.46) 103 (0.88) 9 (2.69) 342 (2.57) 155 (0.78)
0.1 5.7 0.2 <0.1 0.6 0.9
immature Pelecypoda 110 (1.80) 1,263 (2.72) 562 (1.28) 33 (2.74) --- ---
0.3 9.2 1.0 0.1
Arthropoda
Crustacea
Amphipoda (total) 71 (1.60) 42 (2.01) 174 (1.78) 966 (2.61) 5,627 (1.17) 97 (0.69)
0.2 0.3 0.3 3.3 9.4 0.5
Gammaridae
Gammarus sp. <1 (3.61) 1 (6.25) 40 (2.70) 2 (3.06) 2,905 (2.06) 68 (0.65)
<0.1 <0.1 0.1 <0.1 4.9 0.4
127
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv)
Taxon % % % % % %
Hyalellidae
Hyalella azteca 33 (1.74) 34 (2.22) 103 (1.55) 903 (2.64) 2,196 (1.80) ---
0.1 0.2 0.2 3.1 3.7
Decapoda
Cambaridae
Procambarus sp. imm. --- --- 2 (3.07) 1 (5.48) --- ---
<0.1 <0.1
128
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Aquatic Acari (total) 1,075 (0.82) 330 (1.58) 5,560 (1.05) 2,367 (0.90) 1,711 (1.15) 319 (0.74)
2.8 2.4 10.1 8.0 2.9 1.8
Hydracarina 659 (0.71) 268 (1.66) 4,168 (1.17) 1,693 (0.74) 1,341 (1.05) 280 (0.96)
1.7 2.0 7.5 5.8 2.2 1.6
Oribatidae 416 (1.78) 61 (1.87) 1,432 (2.08) 519 (1.82) 369 (1.67) 39 (0.87)
1.1 0.4 2.6 1.8 0.6 0.2
Insecta
Collembola --- 3 (4.61) 1 (4.24) <1 (5.48) 8 (3.87) ---
<0.1 <0.1 <0.1 <0.1
Ephemeroptera (total) 788 (1.19) 48 (2.46) 3,104 (1.03) 1,215 (0.80) 2,596 (1.01) 261 (0.62)
2.0 0.3 5.6 4.1 4.3 1.5
Eurylophella doris --- --- --- --- 8 (3.87) ---
<0.1
Heptageniidae (e.i. or dam.) 2 (3.61) 1 (6.25) 43 (1.34) 254 (1.26) 35 (2.66) 39 (0.87)
<0.1 <0.1 0.1 0.9 0.1 0.2
129
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Leptohyphidae (e.i.) 59 (2.01) --- 300 (1.80) 136 (2.47) 393 (1.67) 97 (1.73)
0.2 0.5 0.5 0.7 0.5
Asioplax dolani --- --- --- 1 (5.48) --- ---
<0.1
Tricorythodes albilineatus 227 (1.62) 8 (4.76) 1,772 (1.32) 280 (1.66) 723 (2.03) ---
0.6 0.1 3.2 1.0 1.2
Odonata
Anisoptera (total) 3 (2.58) 8 (2.52) 5 (2.67) 8 (2.13) 66 (1.84) 19 (1.73)
<0.1 0.1 <0.1 <0.1 0.1 0.1
130
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv)
Taxon % % % % % %
Corduliidae
Neurocordulia sp. --- --- --- 4 (3.02) --- ---
<0.1
Macromiidae
Macromia sp. --- --- --- --- 27 (2.35) ---
<0.1
131
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Mesoveliidae
Mesovelia amoena 3 (2.58) --- --- --- --- ---
<0.1
Trichoptera (total) 2,000 (0.94) 165 (3.18) 2,916 (0.95) 3,691 (1.16) 7,542 (0.77) 431 (0.37)
5.1 1.2 5.3 12.5 12.6 2.4
Hydropsychidae (total) 323 (1.31) 14 (4.30) 461 (1.84) 968 (1.30) 63 (3.52) 29 (1.31)
0.8 0.1 0.8 3.3 0.1 0.2
Hydropsychidae e.i. 67 (1.66) --- 254 (2.19) 308 (1.55) 17 (3.87) 10 (1.73)
0.2 0.5 1.0 <0.1 0.1
Hydropsyche sp. 103 (1.97) 1 (6.25) 62 (1.70) 182 (1.71) 7 (3.87) 5 (1.73)
0.3 <0.1 0.1 0.6 <0.1 <0.1
Hydropsyche rossi 110 (1.41) 9 (5.51) 75 (1.67) 346 (1.26) --- ---
Hydroptilidae (total) 1386 (1.02) 74 (3.21) 2,061 (1.01) 1,291 (0.90) 4,422 (0.91) 193 (0.96)
3.6 0.5 3.7 4.4 7.4 1.1
134
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Hydroptilidae e.i. 202 (0.99) 64 (3.75) 169 (0.99) 222 (1.25) 1,505 (1.39) 39 (1.73)
0.5 0.5 0.3 0.8 2.5 0.2
Hydroptila sp. 1192 (1.06) 49 (3.42) 1,566 (1.12) 1,000 (1.15) 1,748 (0.83) 39 (0.87)
3.1 0.4 2.8 3.4 2.9 0.2
Neotrichia sp. <1 (3.61) 16 (5.81) 127 (3.17) 56 (2.57) 1,078 (1.10) 116 (1.00)
<0.1 0.1 0.2 0.2 1.8 0.7
Leptoceridae (total) 27 (1.55) 72 (5.16) 108 (1.86) 215 (4.15) 1,687 (1.45) 140 (0.69)
0.1 0.5 0.2 0.7 2.8 0.8
Nectopsyche sp. <1 (3.61) 2 (6.25) 7 (1.78) 3 (3.40) 759 (1.51) 48 (1.25)
<0.1 <0.1 <0.1 <0.1 1.3 0.3
unidentifiable imm. Trichoptera 235 (1.47) 6 (3.20) 226 (1.42) 983 (2.44) 1,367 (1.30) 68 (1.73)
0.6 <0.1 0.4 3.3 2.3 0.4
Sisyridae
Sisyra sp. --- --- --- --- 4 (3.87) ---
<0.1
Coleoptera (total) 4,914 (1.67) 239 (1.17) 5,756 (1.24) 3,049 (0.80) 3,025 (1.05) 406 (0.12)
12.6 1.8 10.4 10.4 5.1 2.3
Elmidae (total) 4,909 (1.67) 236 (1.19) 9,163 (2.03) 3,012 (0.80) 3,011 (1.05) 396 (0.11)
12.6 1.7 16.6 10.2 5.0 2.2
Elimidae (e.i.) 185 (1.95) 16 (3.59) 204 (1.52) 68 (2.06) --- 10 (1.73)
0.5 0.1 0.4 0.2 0.1
Dubiraphia vittata 14 (2.49) 58 (2.00) 54 (1.49) 8 (2.69) 594 (0.92) 32 (0.91)
<0.1 0.4 0.1 <0.1 1.0 0.2
Stenelmis sp. 4,536 (1.77) 162 (1.36) 8,783 (2.08) 2,904 (0.76) 2,299 (1.24) 358 (0.17)
11.8 1.2 15.9 9.9 3.8 2.0
Stenelmis hungerfordi or
lignicola 6 (3.08) --- 1 (2.91) 1 (5.48) --- ---
<0.1 <0.1 <0.1
Haliplidae
Peltodytes sp. --- 2 (6.25) 5 (2.91) 1 (5.48) --- ---
<0.1 <0.1 <0.1
139
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Ceratopogonidae (total) 1,533 (1.58) 683 (1.86) 402 (0.83) 632 (1.04) 476 (1.07) 48 (0.35)
3.9 5.0 0.7 2.1 0.8 0.3
Bezzia or Palpomyia 917 (1.51) 230 (2.99) 340 (0.89) 508 (1.13) 375 (1.06) 48 (0.35)
2.4 1.7 0.6 1.7 0.6 0.3
Probezzia sp. --- 293 (2.27) 8 (2.77) <1 (5.48) 19 (2.64) ---
2.1 <0.1 <0.1 <0.1
Ceratopogonidae e.i. 565 (2.08) 132 (1.90) 5 (3.30) 94 (2.27) 69 (2.72) ---
1.5 1.0 <0.1 0.3 0.1
140
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Chironomidae (total) 22,137 (0.95) 6,447 (1.04) 28,049 (1.05) 15,361 (0.73) 31,720 (0.57) 15,167 (0.58)
57.0 47.2 50.8 52.2 53.0 85.8
141
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv) no. m-2 (cv)
Taxon % % % % % %
Diamesinae
Potthastia longimana --- --- --- 13 (3.12) --- ---
<0.1
Orthocladiinae e.i. 242 (1.10) 51 (2.23) 432 (1.61) 89 (2.46) 509 (1.48) 155 (0.57)
0.6 0.4 0.8 0.3 0.9 0.9
Corynoneura sp. --- 31 (1.82) --- 21 (2.05) 289 (1.35) 928 (0.87)
0.2 0.1 0.5 5.2
143
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Corynoneura species B 99 (2.51) 26 (2.09) 43 (2.31) 50 (3.06) 592 (1.29) 8,884 (0.87)
0.3 0.2 0.1 0.2 1.0 50.2
Corynoneura lobata or sp. G --- --- 15 (2.91) --- 59 (3.87) 309 (1.10)
<0.1 0.1 1.7
Cricotopus sp. 353 (2.56) 18 (4.74) 172 (2.23) 106 (1.85) 1,119 (2.55) 19 (1.73)
0.9 0.1 0.3 0.4 1.9 0.1
Cricotopus albiforceps 23 (3.61) 3.41 (6.25) 34 (4.24) 60 (2.73 ) 195 (1.59) 97 (0.92)
0.1 <0.1 0.1 0.2 0.3 0.5
Cricotopus bicinctus 688 (2.43) 33 (4.73) 853 (1.80) 228 (1.92) 4,673 (1.05) 174 (1.45)
1.8 0.2 1.5 0.8 7.8 1.0
144
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Cricotopus albiforceps or
vierriensis 77 (2.28) --- 40 (3.00) 48 (2.90) 40 (2.69) ---
0.2 0.1 0.2 0.1
Cricotopus or Orthocladius 177 (3.20) 24 (4.24) 28 (3.22) 54 (2.64) 643 (3.82) ---
0.5 0.2 0.1 0.2 1.1
Nanocladius crassicornus or
spiniplenus --- --- --- 7 (5.48) --- ---
<0.1
145
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Parakiefferiella sp. 1,680 (1.06) 91 (2.08) 1,015 (0.88) 1,019 (0.84) 1,505 (0.73) 251 (0.74)
4.3 0.7 1.8 3.5 2.5 1.4
Parakiefferiella species E 577 (1.80) 46 (3.42) 116 (2.08) 259 (2.19) 1,300 (1.39) 271 (0.89)
1.5 0.3 0.2 0.9 2.2 1.5
Thienemanniella sp. 315 (1.22) 7 (4.61) 116 (1.39) 68 (2.37) 361 (1.46) 145 (0.92)
0.8 0.1 0.2 0.2 0.6 0.8
146
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Thienemanniella lobapodema 198 (2.21) --- 34 (3.54) 186 (1.76) 535 (1.35) ---
0.5 0.1 0.6 0.9
Thienemanniella species B 1,023 (1.40) 17 (4.83) 633 (1.69) 446 (1.64) 366 (1.47) 77 (1.15)
2.6 0.1 1.1 1.5 0.6 0.4
Tvetenia vitracies 1,229 (2.02) 111 (3.99) 393 (2.07) 966 (1.71) 67 (2.45) ---
3.2 0.8 0.7 3.3 0.1
Chironominae e.i. 146 (2.04) 75 (2.28) 51 (3.14) 25 (2.64) 132 (2.04) ---
0.4 0.6 0.1 0.1 0.2
Cryptotendipes sp. 3 (3.61) 139 (2.97) 196 (3.14) 15 (2.65) 73 (2.66) ---
<0.1 1 0.4 0.1 0.1
Dicrotendipes neomodestus 1,123 (2.05) 79 (4.01) 149 (1.82) 214 (1.68) 519 (1.59) 39 (1.73)
2.9 0.6 0.3 0.7 0.9 0.2
Paralauterborniella
nigrohalteralis --- 11 (5.05) --- --- --- ---
0.1
Polypedilum sp. 1,390 (2.15) 764 (2.11) 2,707 (1.89) 351 (1.45) 232 (3.27) 1,305 (0.73)
3.6 5.6 4.9 1.2 0.4 7.4
Polypedilum flavum 140 (2.27) 24 (3.011) 62 (2.09) 79 (1.70) 247 (1.49) ---
0.4 0.2 0.1 0.3 0.4
Polypedilum scalaenum group 2 (3.61) 464 (2.98) 57 (2.98) 70 (2.35) 454 (2.65) 454 (0.46)
<0.1 3.4 0.1 0.2 0.8 2.6
Stenochironomus sp. 1 (3.61) 16 (4.40) --- 1,046 (1.17) 20 (3.87) 116 (1.73)
<0.1 0.1 3.6 <0.1 0.7
Xestochironomus sp. --- 10 (5.58) --- 1,520 (1.24) 20 (3.87) 435 (0.98)
0.1 5.2 <0.1 2.5
Pseudochironomini
Pseudochironomus sp. 7,537 (1.37) 476 (2.26) 4,965 (1.49) 3,068 (1.92) 296 (1.92) 135 (0.89)
19.4 3.5 9.0 10.4 0.5 0.8
Cladotanytarsus daviesi 113 (2.49) 544 (2.42) 183 (3.07) 42 (2.96) 445 (3.06) 19 (1.73)
0.3 4.0 0.3 0.1 0.7 0.1
Rheotanytarsus exiguus 299 (2.00) 1 (6.25) 327 (2.68) 296 (1.81) 377 (1.27) 19 (1.73)
0.8 <0.1 0.6 1.0 0.6 0.1
Stempellina species A 122 (3.61) 855 (1.78) 2,073 (2.71) 72 (2.03) 542 (2.06) 19 (1.73)
0.3 6.3 3.8 0.2 0.9 0.1
Tanytarsus sp. 590 (2.09) 48 (1.70) 376 (1.50) 187 (1.67) 1,598 (0.82) 77 (1.73)
1.5 0.3 0.7 0.6 2.7 0.4
Tanytarsus species C 3,362 (1.61) 465 (2.14) 11,942 (1.41) 3,252 (1.63) 9,324 (1.32) 232 (1.0)
8.7 3.4 21.6 11.1 15.6 1.3
Tanytarsus species L 2 (3.61) 39 (3.34) 30 (4.24) 6 (5.48) 223 (1.39) 174 (.33)
<0.1 0.3 0.1 <0.1 0.4 1.0
Simuliidae (total) 701 (3.59) 9 (2.29) 472 (3.61) 614 (1.25) 24 (1.85) 29 (1.00)
1.8 0.1 0.9 2.1 <0.1 0.2
154
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Simulium sp. 701 (3.59) 9 (2.29) 469 (3.64) 614 (1.25) 24 (1.85) 29 (1.00)
1.8 0.1 0.8 2.1 <0.1 0.2
155
APPENDIX 1 (cont.)
Habitat
Bedrock Sand Cobble/Gravel Snags Tree Roots Debris Packs
-2 -2 -2 -2 -2
no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m (cv) no. m-2 (cv)
Taxon % % % % % %
Mean Total Organisms 38,865 (0.67) 13,665 (0.84) 55,241 (0.88) 29,413 (0.50) 59,802 (0.59) 17,686 (0.47)
100 100 100 100 100 100
Mean Taxa Richness 27.3 (0.26) 19.9 (0.53) 38.3 (0.10) 35.5 (0.23) 35.0 (0.41) 37.0 (0.08)
Mean Diversity 3.07 (0.14) 2.75 (0.33) 3.49 (0.09) 3.77 (0.16) 3.77 (0.20) 2.97 (0.36)
Mean Eveness 0.65 (0.10) 0.66 (0.19) 0.66 (0.09) 0.74 (0.11) 0.76 (0.11) 0.57 (0.36)
156
APPENDIX 2. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from areally dominant habitats in the Withlacoochee River
study area during 2005-2007.
157
APPENDIX 2A. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the bedrock habitat in the Withlacoochee River study area,
2005-2007.
158
APPENDIX 2B. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the cobble/gravel habitat in the Withlacoochee River study area,
2005-2007.
159
APPENDIX 2C. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the sand habitat in the Withlacoochee River study area, 2005-
2007.
160
APPENDIX 2D. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the snag (wood debris) habitat in the Withlacoochee River study
area, 2005-2007.
161
APPENDIX 2E. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the tree root system habitat in the Withlacoochee River study
area, 2005-2007.
162
APPENDIX 2F. Density (no. m-2 ) and percent composition (%) of the ten most abundant
invertebrate taxa collected from the organic debris pack habitat in the Withlacoochee River study
area, 2005-2007.
163
APPENDIX 3. Habitat suitability curves produced for selected descriptors of invertebrate
community quality and indicator invertebrate taxa using the methods of Gore and Judy 1981.
164
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Taxa Richness collected in the
northern Withlacoochee River, Florida, 2005-2007.
165
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Shannon Diversity collected in the
northern Withlacoochee River, Florida, 2005-2007.
166
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168 183 196
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Total Organisms collected in the
northern Withlacoochee River, Florida, 2005-2007.
167
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity cm
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Hydracarina (Acari) collected in the
northern Withlacoochee River, Florida, 2005-2007.
168
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168 183 196
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Total Baetidae collected in the
northern Withlacoochee River, Florida, 2005-2007.
169
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Pseudocloeon ephippiatum,
(Ephemeroptera:Baetidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
170
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Total Hydropsychidae (Trichoptera)
collected in the northern Withlacoochee River, Florida, 2005-2007.
171
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Hydropsyche sp.
(Trichoptera:Hydropsychidae) collected in the northern Withlacoochee River, Florida, 2005-
2007.
172
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Hydroptila sp.
(Trichoptera:Hydroptilidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
173
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Corydalus cornutus
(Megaloptera:Corydalidae) collected in the northern Withlacoochee Rive r, Florida, 2005-2007.
174
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Stenelmis sp. (Coleoptera:Elmidae)
collected in the northern Withlacoochee River, Florida, 2005-2007.
175
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122 152
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168 183 196 213
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Total Chironomidae
(Insecta:Diptera) collected in the northern Withlacoochee River, Florida, 2005-2007.
176
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Cricotopus bicinctus
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
177
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122 152 183 213
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168 183 196 213 229 244 257 274
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Parakiefferiella sp.
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
178
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91 122
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth(cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Tvetenia vitracies
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
179
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Pseudochironomus sp.
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
180
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43
Velocity (cm)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Stempellina sp. A
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
181
1
0.8
0.6
0.4
0.2
0
0 6 12 18 24 30 37 43 49 55 61 67 73 79 85 91
Velocity (cm/sec)
0.8
Suitability
0.6
0.4
0.2
0
15 30 46 61 76 91 107 122 137 152 168 183 196
Depth (cm)
0.8
0.6
0.4
0.2
0
Bedrock Sand Cobble/Gravel Roots Snags Debris Pack
Substrate
Current velocity, depth, and substrate suitability curves for Tanytarsus sp. C
(Diptera:Chironomidae) collected in the northern Withlacoochee River, Florida, 2005-2007.
182
APPENDIX 4. Results of PHABSIM analysis (weighted usable areas) of selected aquatic
invertebrate descriptors of community quality and indicator taxa.
183
Northern Withlac ooc hee R iver - R eac h 1 - Mac roinvertebrate
C ommunities
140000
120000
100000
WUA (s q ft / 1000 linear ft)
80000
T axa R ichnes s
S hannon
60000 D ivers ity
T otal O rganis ms
40000
20000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
184
Northern Withlac ooc hee R iver - R eac h 1 - Mites
60000
50000
WUA (s q ft / 1000 linear ft)
40000
30000
H ydracarina
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
185
Northern Withlac ooc hee R iver - R eac h 1 - E phemeroptera
60000
50000
WUA (s q ft / 1000 linear fett
40000
30000
T otal B aetidae
P s eudocloeon
ephippiatum
20000
10000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
186
Northern Withlac ooc hee R iver - R eac h 1 - Meg aloptera
20000
18000
16000
14000
WUA (s q ft / 1000 linear ft
12000
10000
C orydalus
8000 cornutus
6000
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
187
Northern Withlac ooc hee R iver - R eac h 1 - T ric hoptera
80000
70000
60000
WUA (s q ft / 1000 linear ft)
50000
40000 T otal
H ydrops ychidae
H ydrops yche s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
188
Northern Withlac ooc hee R iver - R eac h 1 - C oleoptera
60000
50000
WUA (s q ft / 1000 linear ft)
40000
30000
S tenelmis
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
189
Northern Withlac ooc hee R iver - R eac h 1 - Diptera I
120000
100000
WUA (s q ft / 1000 linear ft
80000
T otal
60000 C hironomidae
C ricotopus
bicinctus
T anytars us s p. C
40000 P arakiefferiella s p.
20000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
190
Northern Withlac ooc hee R iver - R eac h 1 - Diptera II
120000
100000
WUA (s q ft / 1000 linear ft)
80000
T otal C hironomidae
60000
P s eudochironomus
s p.
S tempellina s p A.
T vetenia vitracies
40000
20000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
191
Northern Withlac ooc hee R iver - R eac h 2 - Mac roinvertebrate
C ommunities
120000
100000
WUA (s q ft / 1000 linear ft)
80000
S hannon
D ivers ity
T otal O rganis ms
40000
20000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
192
Northern Withlac ooc hee R iver - R eac h 2 - Mites
70000
60000
50000
WUA (s q ft / 1000 linear ft)
40000
H ydracarina
30000
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
193
Northern Withlac ooc hee R iver - R eac h 2 - E phemeroptera
70000
60000
50000
WUA (s q ft / 1000 linear fett
40000
T otal B aetidae
30000 P s eudocloeon
ephippiatum
20000
10000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
194
Northern Withlac ooc hee R iver - R eac h 2 - Meg aloptera
45000
40000
35000
WUA (s q ft / 1000 linear ft
30000
25000
20000 C orydalus
cornutus
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
195
Northern Withlac ooc hee R iver - R eac h 2 - T ric hoptera
70000
60000
50000
WUA (s q ft / 1000 linear ft)
40000
T otal
H ydrops ychidae
30000 H ydrops yche s pp.
H ydroptila s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
196
Northern Withlac ooc hee R iver - R eac h 2 - C oleoptera
60000
50000
WUA (s q ft / 1000 linear ft)
40000
30000
S tenelmis
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
197
Northern Withlac ooc hee R iver - R eac h 2 - Diptera I
100000
90000
80000
70000
WUA (s q ft / 1000 linear ft
60000
T otal
50000 C hironomidae
C ricotopus
bicinctus
40000 T anytars us s p. C
P arakiefferiella s p.
30000
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
198
Northern Withlac ooc hee R iver - R eac h 2- Diptera II
100000
90000
80000
70000
WUA (s q ft / 1000 linear ft)
60000
T otal C hironomidae
50000
P s eudochironomus
s p.
40000 S tempellina s p A.
T vetenia vitracies
30000
20000
10000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
199
Northern Withlac ooc hee R iver - R eac h 3 - Mac roinvertebrate C ommunities
45000
40000
35000
T axa R ichnes s
WUA (s q ft / 1000 linear ft)
S hannon
30000
D ivers ity
T otal O rganis ms
25000
20000
15000
10000
5000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
200
Northern Withlac ooc hee R iver - R eac h 3 - Mites
30000
25000
WUA (s q ft / 1000 linear ft)
20000
Hydracarina
s pp.
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
201
Northern Withlac ooc hee R iver - R eac h 3 - E phemeroptera
30000
25000
WUA (s q ft / 1000 linear fett
T otal B aetidae
20000
P s eudocloeon
ephippiatum
15000
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
202
Northern Withlac ooc hee R iver - R eac h 3 - T ric hoptera
25000
20000
WUA (s q ft / 1000 linear ft)
15000
T otal
Hydrops ychidae
Hydrops yche s pp.
10000
Hydroptila s pp.
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
203
Northern Withlac ooc hee R iver - R eac h 3 - Meg aloptera
18000
16000
14000
C orydalus
WUA (s q ft / 1000 linear ft
12000
cornutus
10000
8000
6000
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
204
Northern Withlac ooc hee R iver - R eac h 3 - C oleoptera
30000
25000
WUA (s q ft / 1000 linear ft)
20000 S tenelmis
s pp.
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
205
Northern Withlac ooc hee R iver - R eac h 3 - Diptera I
35000
30000
25000
WUA (s q ft / 1000 linear ft
20000
T otal
C hironomidae
C ricotopus
15000 bicinctus
T anytars us s p. C
P arakiefferiella s p.
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
206
Northern Withlac ooc hee R iver - R eac h 3 - Diptera II
35000
30000
25000
WUA (s q ft / 1000 linear ft)
20000
T otal C hironomidae
P s eudochironomus
15000 s p.
S tempellina s p A.
T vetenia vitracies
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
207
Northern Withlac ooc hee R iver - R eac h 3.5 - Mac roinvertebrate C ommunities
35000
30000
25000
WUA (s q ft / 1000 linear ft)
20000
T axa R ichnes s
S hannon
15000 D ivers ity
T otal O rganis ms
10000
5000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
208
Northern Withlac ooc hee R iver - R eac h 3.5 - Mites
8000
7000
6000
WUA (s q ft / 1000 linear ft)
5000 Hydracarina
s pp.
4000
3000
2000
1000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
209
Northern Withlac ooc hee R iver - R eac h 3.5 - E phemeroptera
2500
2000
WUA (s q ft / 1000 linear fett
T otal B aetidae
1500
P s eudocloeon
ephippiatum
1000
500
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
210
Northern Withlac ooc hee R iver - R eac h 3.5 - T ric hoptera
3500
3000
2500
WUA (s q ft / 1000 linear ft)
2000
T otal
Hydrops ychidae
1500 Hydrops yche s pp.
Hydroptila s pp.
1000
500
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
211
Northern Withlac ooc hee R iver - R eac h 3.5 - Meg aloptera
700
600
500
WUA (s q ft / 1000 linear ft
400
C orydalus
300
cornutus
200
100
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
212
Northern Withlac ooc hee R iver - R eac h 3.5 - C oleoptera
10000
9000
8000
7000
WUA (s q ft / 1000 linear ft)
6000
5000
S tenelmis
4000 s pp.
3000
2000
1000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
213
Northern Withlac ooc hee R iver - R eac h 3.5 - Diptera I
14000
12000
10000
WUA (s q ft / 1000 linear ft
8000
T otal
C hironomidae
C ricotopus
6000 bicinctus
T anytars us s p. C
P arakiefferiella s p.
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
214
Northern Withlac ooc hee R iver - R eac h 3.5 - Diptera II
14000
12000
10000
WUA (s q ft / 1000 linear ft)
8000
T otal C hironomidae
P s eudochironomus
6000 s p.
S tempellina s p A.
T vetenia vitracies
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
215
Northern Withlac ooc hee R iver - R eac h 4 - Mac roinvertebrate C ommunities
120000
100000
WUA (s q ft / 1000 linear ft)
80000
20000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
216
Northern Withlac ooc hee R iver - R eac h 4 - Mites
35000
30000
25000
Hydracarina
WUA (s q ft / 1000 linear ft)
s pp.
20000
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
217
Northern Withlac ooc hee R iver - R eac h 4 - E phemeroptera
40000
35000
P s eudocloeon
ephippiatum
25000
20000
15000
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
218
Northern Withlac ooc hee R iver - R eac h 4 - T ric hoptera
60000
50000
WUA (s q ft / 1000 linear ft)
40000
30000 T otal
Hydrops ychidae
Hydrops yche s pp.
Hydroptila s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
219
Northern Withlac ooc hee R iver - R eac h 4 - Meg aloptera
10000
9000
8000
C orydalus
7000 cornutus
WUA (s q ft / 1000 linear ft
6000
5000
4000
3000
2000
1000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
220
Northern Withlac ooc hee R iver - R eac h 4 - C oleoptera
45000
40000
35000
WUA (s q ft / 1000 linear ft)
30000
25000
20000 S tenelmis
s pp.
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
221
Northern Withlac ooc hee R iver - R eac h 4 - Diptera I
80000
70000
60000
WUA (s q ft / 1000 linear ft
50000
T otal
40000 C hironomidae
C ricotopus
bicinctus
T anytars us s p. C
30000
P arakiefferiella s p.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
222
Northern Withlac ooc hee R iver - R eac h 4 - Diptera II
80000
70000
60000
WUA (s q ft / 1000 linear ft)
50000
T otal C hironomidae
40000
P s eudochironomus
s p.
S tempellina s p A.
30000
T vetenia vitracies
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
223
Northern Withlac ooc hee R iver - R eac h 5 - Mac roinvertebrate
C ommunities
100000
90000
80000
70000
WUA (s q ft / 1000 linear ft)
60000
S hannon
40000 D ivers ity
T otal O rganis ms
30000
20000
10000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
224
Northern Withlac ooc hee R iver - R eac h 5 - Mites
70000
60000
50000
WUA (s q ft / 1000 linear ft)
40000
H ydracarina
30000
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
225
Northern Withlac ooc hee R iver - R eac h 5 - E phemeroptera
50000
45000
40000
35000
WUA (s q ft / 1000 linear fett
30000
25000
T otal B aetidae
P s eudocloeon
20000
ephippiatum
15000
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
226
Northern Withlac ooc hee R iver - R eac h 5 - Meg aloptera
30000
25000
WUA (s q ft / 1000 linear ft
20000
15000
C orydalus
cornutus
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
227
Northern Withlac ooc hee R iver - R eac h 5 - T ric hoptera
50000
45000
40000
35000
WUA (s q ft / 1000 linear ft)
30000
25000 T otal
H ydrops ychidae
H ydrops yche s pp.
20000
H ydroptila s pp.
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
228
Northern Withlac ooc hee R iver - R eac h 5 - C oleoptera
70000
60000
50000
WUA (s q ft / 1000 linear ft)
40000
S tenelmis
30000
s pp.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
229
Northern Withlac ooc hee R iver - R eac h 5 - Diptera I
80000
70000
60000
WUA (s q ft / 1000 linear ft
50000
T otal
40000 C hironomidae
C ricotopus
bicinctus
T anytars us s p. C
30000
P arakiefferiella s p.
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
230
Northern Withlac ooc hee R iver - R eac h 5 - Diptera II
70000
60000
50000
WUA (s q ft / 1000 linear ft)
40000
T otal C hironomidae
P s eudochironomus
30000 s p.
S tempellina s p A.
T vetenia vitracies
20000
10000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
231
Northern Withlac ooc hee R iver - R eac h 6 - Mac roinvertebrate
C ommunities
35000
30000
25000
WUA (s q ft / 1000 linear ft)
20000
T axa R ichnes s
S hannon
15000 D ivers ity
T otal O rganis ms
10000
5000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
232
Northern Withlac ooc hee R iver - R eac h 6 - Mites
4500
4000
3500
WUA (s q ft / 1000 linear ft)
3000
2500
2000 H ydracarina
s pp.
1500
1000
500
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
233
Northern Withlac ooc hee R iver - R eac h 6 - E phemeroptera
6000
5000
WUA (s q ft / 1000 linear fett
4000
3000
T otal B aetidae
P s eudocloeon
ephippiatum
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
234
Northern Withlac ooc hee R iver - R eac h 6 - Meg aloptera
400
350
300
WUA (s q ft / 1000 linear ft
250
200
C orydalus
cornutus
150
100
50
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
235
Northern Withlac ooc hee R iver - R eac h 6 - T ric hoptera
9000
8000
7000
WUA (s q ft / 1000 linear ft)
6000
5000
T otal
H ydrops ychidae
4000 H ydrops yche s pp.
H ydroptila s pp.
3000
2000
1000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
236
Northern Withlac ooc hee R iver - R eac h 6 - C oleoptera
10000
9000
8000
7000
WUA (s q ft / 1000 linear ft)
6000
5000
S tenelmis
4000 s pp.
3000
2000
1000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
237
Northern Withlac ooc hee R iver - R eac h 6 - Diptera I
18000
16000
14000
WUA (s q ft / 1000 linear ft
12000
10000
T otal
C hironomidae
C ricotopus
8000
bicinctus
T anytars us s p. C
6000 P arakiefferiella s p.
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
238
Northern Withlac ooc hee R iver - R eac h 6 - Diptera II
16000
14000
12000
WUA (s q ft / 1000 linear ft)
10000
T otal C hironomidae
8000
P s eudochironomus
s p.
S tempellina s p A.
6000
T vetenia vitracies
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
239
Northern Withlac ooc hee R iver - R eac h 7 - Mac roinvertebrate
C ommunities
90000
80000
70000
WUA (s q ft / 1000 linear ft)
60000
50000
T axa R ichnes s
40000 S hannon
D ivers ity
T otal O rganis ms
30000
20000
10000
0
135 150 165 190 220 250 280 310 340 370 400 475 550 650 800 950 1100 1300 2500
Dis c harg e (c fs )
240
Northern Withlac ooc hee R iver - R eac h 7 - Mites
16000
14000
12000
WUA (s q ft / 1000 linear ft)
10000
8000
H ydracarina
s pp.
6000
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
241
Northern Withlac ooc hee R iver - R eac h 7 - E phemeroptera
14000
12000
10000
WUA (s q ft / 1000 linear fett
8000
T otal B aetidae
6000 P s eudocloeon
ephippiatum
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
1500
2500
Dis c harg e (c fs )
242
Northern Withlac ooc hee R iver - R eac h 7 - Meg aloptera
1200
1000
WUA (s q ft / 1000 linear ft
800
600
C orydalus
cornutus
400
200
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
243
Northern Withlac ooc hee R iver - R eac h 7 - T ric hoptera
25000
20000
WUA (s q ft / 1000 linear ft)
15000
T otal
H ydrops ychidae
H ydrops yche s pp.
10000
H ydroptila s pp.
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
244
Northern Withlac ooc hee R iver - R eac h 7 - C oleoptera
18000
16000
14000
WUA (s q ft / 1000 linear ft)
12000
10000
8000 S tenelmis
s pp.
6000
4000
2000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
245
Northern Withlac ooc hee R iver - R eac h 7 - Diptera I
50000
45000
40000
35000
WUA (s q ft / 1000 linear ft
30000
T otal
25000 C hironomidae
C ricotopus
bicinctus
20000 T anytars us s p. C
P arakiefferiella s p.
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
246
Northern Withlac ooc hee R iver - R eac h 7 - Diptera II
50000
45000
40000
35000
WUA (s q ft / 1000 linear ft)
30000
T otal C hironomidae
25000
P s eudochironomus
s p.
20000 S tempellina s p A.
T vetenia vitracies
15000
10000
5000
0
5
5
5
5
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
00
00
00
00
00
13
14
15
16
18
20
22
24
26
28
30
32
34
36
38
40
45
50
55
60
70
80
90
10
11
12
15
25
Dis c harg e (c fs )
247
APPENDIX 5. Time series analyses of aquatic invertebrate community descriptor and indicator
taxa PHABSIM results.
248
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 1
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
April
August
May
July
January
February
November
September
October
December
June
March
249
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 1
120%
100%
80%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
250
Shannon Diversity (1940-1969)
Northern Withlacoochee River - Reach 1
80%
70%
60%
50%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
30%
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
-10%
April
August
May
July
January
February
November
September
October
December
June
March
251
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 1
120%
100%
80%
Habitat GAin / Loss
60%
10% Flow Reduction
20% Flow Reduction
40% 30% Flow Reduction
40% Flow Reduction
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
252
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 1
40%
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
253
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 1
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
254
Hydracarina sp. 1940-1969
Northern Withlacoochee River Reach 1
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
-10%
April
August
May
July
February
January
November
March
June
October
September
December
255
Hydracarina sp. 1970-1999
Northern Withlacoochee River Reach 1
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
256 December
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 1
40%
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
257
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 1
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20% 30% Flow Reduction
40% Flow Reduction
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
258
Pseudocloeon ephippiatum (1940-1969)
Withlacoochee River - Reach 1
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
259
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 1
140%
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
260
Corydalus cornutus 1940-1969
Northern Withlacoochee river Reach 1
120%
100%
80%
Habitat Gain / Loss
60%
40%
10% Flow Reduction
20% Flow Reduction
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
261
Corydalus cornutus 1970-1999
Northern Withlacoochee River Reach 1
180%
160%
140%
120%
100%
Habitat Gain / Loss
20%
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
262 December
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 1
50%
40%
30%
20%
Habitat Gain/Loss
-10%
-20%
-30%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
263
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 1
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
264
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 1
50%
40%
30%
20%
Habitat Gain/Loss
10%
10% Flow Reduction
20% Flow Reduction
0%
30% Flow Reduction
40% Flow Reduction
-10%
-20%
-30%
-40%
-50%
April
August
May
July
February
January
November
March
June
October
September
December
265
Hydropsyche spp. (1970-1999)
Withlacoochee River - Reach 1
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
266
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 1
60%
50%
40%
30%
Habitat Gain / Loss
20%
10%
10% Flow Reduction
20% Flow Reduction
0%
30% Flow Reduction
40% Flow Reduction
-10%
-20%
-30%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
267
Hydroptila sp. 1970-1999
Northern Withlacoochee River Reach 1
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
268 December
Stenelmis sp. 1940-1969
Northern Withlacoochee River Reach 1
120%
100%
80%
Habitat Gain / Loss
60%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% 40% Flow Reduction
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
269
Stenelmis sp. 1970-1999
Northern Withlacoochee River Reach 1
160%
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
270 December
Total Chironomidae 1940-1969
Northern Withlacoochee River - Reach 1
30%
20%
10%
Habitat G ain / L os s
0%
10% F low
R eduction
-10% 20% F low
R eduction
30% F low
R eduction
-20% 40% F low
R eduction
-30%
-40% A ug us t
A pril
November
Oc tober
Dec ember
S eptember
J une
J uly
J anuary
F ebruary
May
Marc h
271
Total Chironomidae 1970-1999
Northern Withlacoochee River Reach 1
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
272 December
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River Reach 1
0%
-5%
Habitat Gain / Loss
-10%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
-15%
-20%
-25%
April
August
May
July
February
January
November
March
June
October
September
December
273
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River Reach 1
0%
-5%
Habitat Gain / Loss
-10%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
-15%
-20%
-25%
April
August
May
July
February
January
November
March
June
October
September
274 December
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River Reach 1
40%
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
275
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River Reach 1
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
276 December
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River Reach 1
15%
10%
5%
0%
Habitat Gain / Loss
-5%
10% Flow Reduction
20% Flow Reduction
-10%
30% Flow Reduction
40% Flow Reduction
-15%
-20%
-25%
-30%
-35%
April
August
May
July
February
January
November
March
June
October
September
December
277
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River Reach 1
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
-40%
April
August
May
July
February
January
November
March
June
October
September
278 December
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River Reach 1
30%
20%
10%
0%
-10%
Habitat Gain / Loss
-40%
-50%
-60%
-70%
-80%
April
August
May
July
February
January
November
March
June
October
September
December
279
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River Reach 1
80%
60%
40%
20%
Habitat Gain / Loss
-40%
-60%
-80%
April
August
May
July
February
January
November
March
June
October
September
280 December
Stempellina sp. A 1940-1969
Northern Withlacoochee River Reach 1
600%
500%
400%
Habitat Gain / Loss
200%
100%
0%
April
August
May
July
February
January
November
March
June
October
September
December
281
Stempellina sp. A 1970-1999
Northern Withlacoochee River Reach 1
700%
600%
500%
Habitat Gain / Loss
200%
100%
0%
April
August
May
July
February
January
November
March
June
October
September
282 December
Tvetenia vitracies 1940-1969
Northern Withlacoochee River Reach 1
60%
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
-40%
-50%
April
August
May
July
February
January
November
March
June
October
September
December
283
Tvetenia vitracies 1970-1999
Northern Withlacoochee River Reach 1
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
April
August
May
July
February
January
November
March
June
October
September
284 December
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 2
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
-10%
April
August
May
July
January
February
November
September
October
December
June
March
285
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 2
120%
100%
80%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
286
Shannon Diversity 1940-1969
Northern Withlacoochee River Reach 2
90%
80%
70%
60%
Habitat Gain / Loss
50%
40%
10%
0%
-10%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
287
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 2
140%
120%
100%
80%
Habitat GAin / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
288
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 2
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
January
February
November
September
October
December
June
March
289
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 2
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
290
Hydracarina sp. 1940-1969
Northern Withlacoochee River Reach 2
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
291
Hydracarina sp. 1970-1999
Northern Withlacoochee River Reach 2
180%
160%
140%
120%
Habitat Gain / Loss
60%
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
292 December
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 2
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
January
February
November
September
October
December
June
March
293
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 2
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20% 30% Flow Reduction
40% Flow Reduction
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
294
Pseudocloeon ephippiatum (1940-1969)
Withlacoochee River - Reach 2
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
295
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 2
140%
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
296
Corydalus cornutus 1940-1969
Northern Withlacoochee river Reach 2
140%
120%
100%
Habitat Gain / Loss
80%
60%
10% Flow Reduction
20% Flow Reduction
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
297
Corydalus cornutus 1970-1999
Northern Withlacoochee River Reach 2
200%
180%
160%
140%
Habitat Gain / Loss
120%
10% Flow Reduction
20% Flow Reduction
100%
30% Flow Reduction
40% Flow Reduction
80%
60%
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
298 December
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 2
80%
60%
40%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
299
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 2
120%
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
300
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 2
80%
60%
40%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
301
Hydropsyche spp. (1970-1999)
Withlacoochee River - Reach 2
120%
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
302
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 2
70%
60%
50%
40%
Habitat Gain / Loss
30%
20%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
303
Hydroptila sp. 1970-1999
Northern Withlacoochee River Reach 2
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
304 December
Stenelmis sp. 1940-1969
Northern Withlacoochee River Reach 2
140%
120%
100%
Habitat Gain / Loss
80%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
305
Stenelmis sp. 1970-1999
Northern Withlacoochee River Reach 2
180%
160%
140%
120%
Habitat Gain / Loss
60%
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
306 December
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 2
40%
30%
20%
Habitat G ain / L os s
10%
0% 10% F low
R eduction
20% F low
R eduction
-10% 30% F low
R eduction
40% F low
R eduction
-20%
-30%
-40% A ug us t
A pril
November
Oc tober
Dec ember
S eptember
J une
J uly
J anuary
F ebruary
May
Marc h
307
Total Chironomidae 1970-1999
Northern Withlacoochee River Reach 2
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
308 December
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River Reach 2
20%
15%
10%
5%
Habitat Gain / Loss
-10%
-15%
-20%
-25%
April
August
May
July
February
January
November
March
June
October
September
December
309
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River Reach 2
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
310 December
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River Reach 2
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
311
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River Reach 2
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
312 December
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River Reach 2
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
313
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River Reach 2
80%
60%
40%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
314 December
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River Reach 2
30%
20%
10%
0%
Habitat Gain / Loss
-10%
10% Flow Reduction
20% Flow Reduction
-20%
30% Flow Reduction
40% Flow Reduction
-30%
-40%
-50%
-60%
-70%
April
August
May
July
February
January
November
March
June
October
September
December
315
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River Reach 2
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
-80%
April
August
May
July
February
January
November
March
June
October
September
December
316
Stempellina sp. A 1940-1969
Northern Withlacoochee River Reach 2
600%
500%
400%
Habitat Gain / Loss
200%
100%
0%
April
August
May
July
February
January
November
March
June
October
September
December
317
Stempellina sp. A 1970-1999
Northern Withlacoochee River Reach 2
700%
600%
500%
Habitat Gain / Loss
200%
100%
0%
April
August
May
July
February
January
November
March
June
October
September
318 December
Tvetenia vitracies 1940-1969
Northern Withlacoochee River Reach 2
60%
50%
40%
30%
Habitat Gain / Loss
20%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
-30%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
319
Tvetenia vitracies 1970-1999
Northern Withlacoochee River Reach 2
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
320 December
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 3
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
321
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 3
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
322
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 3
120%
100%
80%
Habitat GAin / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
323
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 3
70%
60%
50%
Habitat Gain / Loss
40%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
324
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 3
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
325
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 3
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
326
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 3
180%
160%
140%
120%
Habitat Gain / Loss
60%
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
327
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 3
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
328
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 3
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
329
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 3
50%
40%
30%
20%
Habitat Gain / Loss
10%
10% Flow Reduction
20% Flow Reduction
0% 30% Flow Reduction
40% Flow Reduction
-10%
-20%
-30%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
330
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 3
200%
150%
Habitat Gain/Loss
100%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
50%
0%
-50%
August
July
April
May
June
October
March
January
February
November
September
December
331
Pseudocloeon ephippiatum (1940-1969)
Withlacoochee River - Reach 3
140%
120%
100%
80%
Habitat Gain/Loss
40%
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
December
September
332
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 3
120%
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
333
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 3
80%
70%
60%
50%
40%
Habitat Gain/Loss
10%
0%
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
334
Hydropsyche spp. (1970-1999)
Northern Withlacoochee River - Reach 3
120%
100%
80%
60%
Habitat Gain/Loss
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
335
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 3
60%
50%
40%
30%
Habitat Gain/Loss
0%
-10%
-20%
-30%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
336
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 3
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
337
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 3
70%
60%
50%
40%
Habitat Gain / Loss
30%
20%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
-30%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
338
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 3
300%
250%
200%
Habitat Gain / Loss
50%
0%
-50%
August
July
October
April
May
June
January
March
November
February
De cem ber
Septem ber
339
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 3
200%
150%
Habitat Gain / Loss
100%
0%
-50%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
340
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 3
180%
160%
140%
120%
Habitat Gain / Loss
60%
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
341
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 3
140%
120%
100%
Habitat Gain / Loss
80%
20%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
342
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 3
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
343
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 3
40%
30%
20%
H abitat G ain / L os s
10%
0% 10% F low
R educ tion
20% F low
R educ tion
-10% 30% F low
R educ tion
40% F low
-20% R educ tion
-30%
-40% A ug us t
A pril
Oc tober
S eptember
November
D ec ember
J une
Marc h
May
J anuary
F ebruary
J uly
344
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 3
30%
25%
20%
15%
Habitat Gain / Loss
10%
10% Flow Reduction
20% Flow Reduction
5%
30% Flow Reduction
40% Flow Reduction
0%
-5%
-10%
-15%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
345
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 3
15%
10%
5%
Habitat Gain / Loss
-10%
-15%
-20%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
346
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 3
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
347
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 3
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
Oct ober
April
January
June
May
Nov ember
March
February
December
Septem ber
348
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 3
70%
60%
50%
40%
Habitat Gain / Loss
30%
10% Flow Reduction
20% Flow Reduction
20%
30% Flow Reduction
40% Flow Reduction
10%
0%
-10%
-20%
-30%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
349
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 3
20%
15%
10%
5%
Habitat Gain / Loss
0%
10% Flow Reduction
20% Flow Reduction
-5%
30% Flow Reduction
40% Flow Reduction
-10%
-15%
-20%
-25%
-30%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
350
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 3
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
-80%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
351
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 3
30%
20%
10%
0%
Habitat Gain / Loss
-10%
10% Flow Reduction
20% Flow Reduction
-20%
30% Flow Reduction
40% Flow Reduction
-30%
-40%
-50%
-60%
-70%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
352
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 3
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
353
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 3
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
354
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 3
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
355
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 3
70%
60%
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
-30%
-40%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
356
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
357
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 3.5
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
358
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat GAin / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
359
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 3.5
70%
60%
50%
Habitat Gain / Loss
40%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
360
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
361
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 3.5
40%
30%
20%
Habitat Gain / Loss
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
362
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
363
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 3.5
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
364
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
365
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 3.5
40%
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
366
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 3.5
250%
200%
150%
Habitat Gain/Loss
0%
-50%
-100%
August
July
April
May
June
October
March
January
February
November
December
September
367
Pseudocloeon ephippiatum (1940-1969)
Northern Withlacoochee River - Reach 3.5
140%
120%
100%
80%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
40%
30% Flow Reduction
40% Flow Reduction
20%
0%
-20%
-40%
-60%
August
July
April
May
June
October
March
January
February
November
September
December
368
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
369
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 3.5
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
370
Hydropsyche spp. (1970-1999)
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
371
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 3.5
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
372
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
373
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 3.5
40%
30%
20%
Habitat Gain / Loss
10%
-20%
-30%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
374
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 3.5
800%
700%
600%
500%
Habitat Gain / Loss
400%
10% Flow Reduction
20% Flow Reduction
300%
30% Flow Reduction
40% Flow Reduction
200%
100%
0%
-100%
-200%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
375
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 3.5
300%
250%
200%
Habitat Gain / Loss
150%
100%
10% Flow Reduction
20% Flow Reduction
0%
-50%
-100%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
376
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
377
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 3.5
80%
70%
60%
Habitat Gain / Loss
50%
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
378
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
379
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 3.5
40%
30%
20%
H abitat G ain / L os s
10%
0% 10% F low
R educ tion
20% F low
R educ tion
-10% 30% F low
R educ tion
40% F low
-20% R educ tion
-30%
-40% A ug us t
A pril
Oc tober
S eptember
November
D ec ember
J une
Marc h
May
J anuary
F ebruary
J uly
380
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 3.5
20%
15%
10%
Habitat Gain / Loss
-5%
-10%
-15%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
381
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 3.5
15%
10%
5%
Habitat Gain / Loss
-5%
-10%
-15%
August
July
Oct ober
April
January
June
May
Nov ember
March
February
December
Septem ber
382
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 3.5
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
383
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 3.5
20%
15%
10%
5%
Habitat Gain / Loss
0%
10% Flow Reduction
20% Flow Reduction
-5%
30% Flow Reduction
40% Flow Reduction
-10%
-15%
-20%
-25%
-30%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
384
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
385
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 3.5
20%
15%
10%
5%
Habitat Gain / Loss
-10%
-15%
-20%
-25%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
386
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
387
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 3.5
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
388
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 3.5
20%
10%
0%
Habitat Gain / Loss
-30%
-40%
-50%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
389
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 3.5
200%
180%
160%
140%
Habitat Gain / Loss
120%
10% Flow Reduction
20% Flow Reduction
100%
30% Flow Reduction
40% Flow Reduction
80%
60%
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
390
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 3.5
250%
200%
Habitat Gain / Loss
150%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
100%
50%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
391
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 3.5
120%
100%
80%
Habitat Gain / Loss
20%
0%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
392
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 3.5
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
-20%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
393
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 4
100%
90%
80%
70%
Habitat Gain / Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
394
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 4
50%
45%
40%
35%
Habitat Gain / Loss
30%
10% Flow Reduction
20% Reduction
25%
30% Flow Reduction
40% Flow Reduction
20%
15%
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
395
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 4
100%
90%
80%
70%
60%
Habitat GAin / Loss
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
396
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 4
60%
50%
40%
Habitat Gain / Loss
30%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
20% 40% Flow Reduction
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
397
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
398
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 4
35%
30%
25%
20%
Habitat Gain / Loss
15%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
5%
0%
-5%
-10%
-15%
August
July
April
May
June
October
March
January
February
November
September
December
399
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 4
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
400
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 4
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
401
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
0%
-20%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
402
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 4
30%
25%
20%
15%
10%
Habitat Gain / Loss
-5%
-10%
-15%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
403
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
404
Pseudocloeon ephippiatum (1940-1969)
Northern Withlacoochee River - Reach 4
50%
40%
30%
Habitat Gain/Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
405
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
406
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 4
50%
40%
30%
Habitat Gain/Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
407
Hydropsyche spp. (1970-1999)
Northern Withlacoochee River - Reach 4
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
408
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 4
60%
50%
40%
Habitat Gain/Loss
10%
0%
-10%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
409
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
410
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 4
35%
30%
25%
20%
Habitat Gain / Loss
15%
10%
10% Flow Reduction
20% Flow Reduction
5%
30% Flow Reduction
40% Flow Reduction
0%
-5%
-10%
-15%
August
July
Oct ober
April
May
June
January
Nov ember
March
February
December
Septem ber
411
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
412
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 4
60%
50%
40%
Habitat Gain / Loss
30%
20%
10% Flow Reduction
20% Flow Reduction
0%
-10%
-20%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
413
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 4
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
414
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 4
80%
70%
60%
Habitat Gain / Loss
50%
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
415
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
416
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 4
30%
25%
20%
15%
H abitat G ain / L os s
10%
5% 10% F low
R educ tion
20% F low
0% R educ tion
30% F low
R educ tion
-5% 40% F low
R educ tion
-10%
-15%
-20% A ug us t
A pril
Oc tober
S eptember
November
D ec ember
J une
Marc h
May
J anuary
F ebruary
J uly
417
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 4
40%
35%
30%
25%
20%
Habitat Gain / Loss
5%
0%
-5%
-10%
-15%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
418
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 4
20%
15%
10%
Habitat Gain / Loss
-5%
-10%
-15%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
419
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
420
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 4
30%
25%
20%
15%
Habitat Gain / Loss
10%
10% Flow Reduction
20% Flow Reduction
5%
30% Flow Reduction
40% Flow Reduction
0%
-5%
-10%
-15%
-20%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
421
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 4
30%
20%
10%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
422
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 4
15%
10%
5%
0%
Habitat Gain / Loss
-15%
-20%
-25%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
423
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 4
30%
20%
10%
0%
Habitat Gain / Loss
-30%
-40%
-50%
-60%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
424
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 4
10%
0%
-10%
Habitat Gain / Loss
-30%
-40%
-50%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
425
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 4
450%
400%
350%
300%
Habitat Gain / Loss
150%
100%
50%
0%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
426
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 4
400%
350%
300%
250%
Habitat Gain / Loss
100%
50%
0%
August
July
Oct ober
April
January
June
May
Nov ember
March
February
December
Septem ber
427
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 4
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
August
July
Oct ober
April
May
January
June
Nov ember
March
February
December
Septem ber
428
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 4
35%
30%
25%
20%
15%
Habitat Gain / Loss
0%
-5%
-10%
-15%
-20%
August
July
Oct ober
April
January
May
June
Nov ember
March
February
December
Septem ber
429
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 5
45%
40%
35%
30%
Habitat Gain / Loss
15%
10%
5%
0%
April
August
May
July
January
February
November
September
October
December
June
March
430
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 5
100%
90%
80%
70%
Habitat Gain / Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
431
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 5
50%
45%
40%
35%
Habitat Gain / Loss
30%
25%
10% Flow Reduction
20% Flow Reduction
20% 30% Flow Reduction
40% Flow Reduction
15%
10%
5%
0%
April
August
May
July
January
February
November
September
October
December
June
March
432
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 5
100%
90%
80%
70%
60%
Habitat GAin / Loss
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
433
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 5
35%
30%
25%
20%
Habitat Gain / Loss
15%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
5%
0%
-5%
-10%
-15%
April
August
May
July
January
February
November
September
October
December
June
March
434
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
435
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 5
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
April
August
May
July
February
January
November
March
June
October
September
December
436
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 5
120%
100%
80%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
437 December
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 5
50%
40%
30%
Habitat Gain / Loss
20%
10% Flow Reduction
20% Flow Reduction
10% 30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
438
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
439
Pseudocloeon ephippiatum (1940-1969)
Northern Withlacoochee River - Reach 5
45%
40%
35%
30%
Habitat Gain/Loss
25%
10% Flow Reduction
20% Flow Reduction
20%
30% Flow Reduction
40% Flow Reduction
15%
10%
5%
0%
-5%
April
August
May
July
January
February
November
September
October
December
June
March
440
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
441
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 5
50%
40%
30%
Habitat Gain / Loss
20%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
10% 40% Flow Reduction
0%
-10%
April
August
May
July
February
January
November
March
June
October
September
December
442
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
443 December
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 5
45%
40%
35%
30%
Habitat Gain/Loss
15%
10%
5%
0%
April
August
May
July
January
February
November
September
October
December
June
March
444
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 5
100%
90%
80%
70%
Habitat Gain/Loss
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
445
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 5
45%
40%
35%
30%
Habitat Gain/Loss
25%
10% Flow Reduction
20% Flow Reduction
20%
30% Flow Reduction
40% Flow Reduction
15%
10%
5%
0%
-5%
April
August
May
July
February
January
November
March
June
October
September
December
446
Hydropsyche spp. (1970-1999)
Northern Withlacoochee River - Reach 5
100%
90%
80%
70%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
447
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 5
40%
35%
30%
25%
Habitat Gain / Loss
20%
15%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
5%
0%
-5%
-10%
April
August
May
July
February
January
November
March
June
October
September
December
448
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
449 December
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 5
80%
70%
60%
Habitat Gain / Loss
50%
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
April
August
May
July
February
January
November
March
June
October
September
December
450
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 5
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
451 December
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 5
40%
30%
20%
Habitat G ain / L os s
-10%
-20% A ug us t
A pril
November
Oc tober
Dec ember
S eptember
J une
J uly
J anuary
F ebruary
May
Marc h
452
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
453 December
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 5
20%
15%
10%
Habitat Gain / Loss
-5%
-10%
-15%
April
August
May
July
February
January
November
March
June
October
September
December
454
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 5
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
April
August
May
July
February
January
November
March
June
October
September
455 December
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 5
25%
20%
15%
10%
Habitat Gain / Loss
-5%
-10%
-15%
April
August
May
July
February
January
November
March
June
October
September
December
456
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 5
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
April
August
May
July
February
January
November
March
June
October
September
457 December
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 5
15%
10%
5%
0%
Habitat Gain / Loss
-15%
-20%
-25%
April
August
May
July
February
January
November
March
June
October
September
December
458
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 5
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
459 December
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 5
20%
10%
0%
-10%
Habitat Gain / Loss
-40%
-50%
-60%
-70%
April
August
May
July
February
January
November
March
June
October
September
December
460
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 5
100%
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
-80%
April
August
May
July
February
January
November
March
June
October
September
December
461
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 5
300%
250%
200%
Habitat Gain / Loss
100%
50%
0%
April
August
May
July
February
January
November
March
June
October
September
December
462
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 5
350%
300%
250%
Habitat Gain / Loss
100%
50%
0%
April
August
May
July
February
January
November
March
June
October
September
463 December
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 5
40%
30%
20%
Habitat Gain / Loss
0%
-10%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
464
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 5
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
465 December
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 6
90%
80%
70%
60%
Habitat Gain / Loss
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
466
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 6
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
467
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 6
90%
80%
70%
60%
Habitat Gain / Loss
50%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
468
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 6
140%
120%
100%
Habitat GAin / Loss
80%
10% Flow Reduction
20% Flow Reduction
60% 30% Flow Reduction
40% Flow Reduction
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
469
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 6
50%
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
January
February
November
September
October
December
June
March
470
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
471
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 6
90%
80%
70%
60%
Habitat Gain / Loss
30%
20%
10%
0%
April
August
May
July
February
January
November
March
June
October
September
December
472
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 6
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
473
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 6
40%
30%
20%
10%
Habitat Gain / Loss
-20%
-30%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
474
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20% 30% Flow Reduction
40% Flow Reduction
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
475
Pseudocloeon ephippiatum (1940-1969)
Withlacoochee River - Reach 6
80%
70%
60%
50%
Habitat Gain/Loss
40%
10% Flow Reduction
20% Flow Reduction
30%
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
-10%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
476
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 6
140%
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
477
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
40%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
478
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 6
160%
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
60%
30% Flow Reduction
40% Flow Reduction
40%
20%
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
479
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 6
80%
70%
60%
50%
Habitat Gain/Loss
40%
10% Flow Reduction
20% Flow Reduction
30%
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
-10%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
480
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 6
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
481
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 6
80%
70%
60%
50%
Habitat Gain/Loss
40%
10% Flow Reduction
20% Flow Reduction
30%
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
-10%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
482
Hydropsyche spp. (1970-1999)
Northern Withlacoochee River - Reach 6
120%
100%
80%
Habitat Gain/Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
483
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 6
60%
50%
40%
30%
Habitat Gain / Loss
20%
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
484
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 6
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
485
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 6
100%
90%
80%
70%
Habitat Gain / Loss
60%
50%
10% Flow Reduction
20% Flow Reduction
40%
30% Flow Reduction
40% Flow Reduction
30%
20%
10%
0%
April
August
May
July
February
January
November
March
June
October
September
December
486
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 6
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
487
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 6
40%
30%
20%
Habitat G ain / L os s
10%
0% 10% F low
R eduction
20% F low
R eduction
-10% 30% F low
R eduction
40% F low
R eduction
-20%
-30%
-40% A ug us t
A pril
November
Oc tober
Dec ember
S eptember
J une
J uly
J anuary
F ebruary
May
Marc h
488
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
489
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 6
4%
2%
0%
-2%
Habitat Gain / Loss
-4%
10% Flow Reduction
20% Flow Reduction
-6%
30% Flow Reduction
40% Flow Reduction
-8%
-10%
-12%
-14%
-16%
April
August
May
July
February
January
November
March
June
October
September
December
490
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 6
10%
5%
Habitat Gain / Loss
0%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
-5%
-10%
-15%
April
August
May
July
February
January
November
March
June
October
September
December
491
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 6
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
492
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
493
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 6
30%
20%
10%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
494
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 6
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
495
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 6
30%
20%
10%
0%
Habitat Gain / Loss
-30%
-40%
-50%
April
August
May
July
February
January
November
March
June
October
September
December
496
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 6
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
April
August
May
July
February
January
November
March
June
October
September
December
497
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 6
300%
250%
200%
Habitat Gain / Loss
50%
0%
-50%
April
August
May
July
February
January
November
March
June
October
September
December
498
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 6
600%
500%
400%
Habitat Gain / Loss
100%
0%
-100%
April
August
May
July
February
January
November
March
June
October
September
December
499
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 6
80%
60%
40%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
500
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 6
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
501
Taxa Richness (1940-1969)
Northern Withlacoochee River - Reach 7
90%
80%
70%
60%
Habitat Gain / Loss
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
502
Taxa Richness (1970-1999)
Northern Withlacoochee River - Reach 7
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
503
Shannon Diversity 1940-1969
Northern Withlacoochee River - Reach 7
100%
90%
80%
70%
Habitat Gain / Loss
60%
50%
10% Flow Reduction
20% Flow Reduction
40% 30% Flow Reduction
40% Flow Reduction
30%
20%
10%
0%
April
August
May
July
January
February
November
September
October
December
June
March
504
Shannon Diversity 1970-1999
Northern Withlacoochee River - Reach 7
140%
120%
100%
Habitat GAin / Loss
80%
10% Flow Reduction
20% Flow Reduction
60% 30% Flow Reduction
40% Flow Reduction
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
505
Total Organisms (1940-1969)
Northern Withlacoochee River - Reach 7
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
506
Total Organisms (1970-1999)
Northern Withlacoochee River - Reach 7
120%
100%
80%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
January
February
November
September
October
December
June
March
507
Hydracarina sp. 1940-1969
Northern Withlacoochee River - Reach 7
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
508
Hydracarina sp. 1970-1999
Northern Withlacoochee River - Reach 7
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
509 December
Total Baetidae (1940-1969)
Northern Withlacoochee River - Reach 7
70%
60%
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
-30%
April
August
May
July
January
February
November
September
October
December
June
March
510
Total Baetidae (1970-1999)
Northern Withlacoochee River - Reach 7
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
January
February
November
September
October
December
June
March
511
Pseudocloeon ephippiatum (1940-1969)
Withlacoochee River - Reach 7
140%
120%
100%
Habitat Gain/Loss
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
512
Pseudocloeon ephippiatum (1970-1999)
Northern Withlacoochee River - Reach 7
250%
200%
Habitat Gain/Loss
150%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
100%
50%
0%
April
August
May
July
January
February
November
September
October
December
June
March
513
Corydalus cornutus 1940-1969
Northern Withlacoochee River - Reach 7
250%
200%
150%
Habitat Gain / Loss
100%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
50% 40% Flow Reduction
0%
-50%
April
August
May
July
February
January
November
March
June
October
September
December
514
Corydalus cornutus 1970-1999
Northern Withlacoochee River - Reach 7
500%
450%
400%
350%
Habitat Gain / Loss
300%
10% Flow Reduction
20% Flow Reduction
250%
30% Flow Reduction
40% Flow Reduction
200%
150%
100%
50%
0%
April
August
May
July
February
January
November
March
June
October
September
515 December
Total Hydropsychidae (1940-1969)
Northern Withlacoochee River - Reach 7
180%
160%
140%
120%
Habitat Gain/Loss
60%
40%
20%
0%
April
August
May
July
January
February
November
September
October
December
June
March
516
Total Hydropsychidae (1970-1999)
Northern Withlacoochee River - Reach 7
300%
250%
200%
Habitat Gain/Loss
100%
50%
0%
April
August
May
July
January
February
November
September
October
December
June
March
517
Hydropsyche spp. (1940-1969)
Northern Withlacoochee River - Reach 7
160%
140%
120%
100%
Habitat Gain/Loss
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
518
Hydropsyche spp. (1970-1999)
Withlacoochee River - Reach 7
250%
200%
Habitat Gain/Loss
150%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
100%
50%
0%
April
August
May
July
January
February
November
September
October
December
June
March
519
Hydroptila sp. - 1940-1969
Northern Withlacoochee River - Reach 7
100%
80%
60%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
20% 40% Flow Reduction
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
520
Hydroptila sp. 1970-1999
Northern Withlacoochee River - Reach 7
160%
140%
120%
100%
Habitat Gain / Loss
40%
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
521
Stenelmis sp. 1940-1969
Northern Withlacoochee River - Reach 7
140%
120%
100%
Habitat Gain / Loss
80%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
December
522
Stenelmis sp. 1970-1999
Northern Withlacoochee River - Reach 7
180%
160%
140%
120%
Habitat Gain / Loss
60%
40%
20%
0%
April
August
May
July
February
January
November
March
June
October
September
523 December
T otal C hironomidae 1940-1969
Northern Withlac ooc hee R iver - R eac h 7
60%
50%
40%
30%
Habitat G ain / L os s
20%
10% F low R eduction
10% 20% F low R eduction
30% F low R eduction
0% 40% F low R eduction
-10%
-20%
-30% A ug us t
A pril
November
Oc tober
Dec ember
S eptember
J une
J uly
J anuary
F ebruary
May
Marc h
524
Total Chironomidae 1970-1999
Northern Withlacoochee River - Reach 7
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
525 December
Cricotopus bicinctus 1940-1969
Northern Withlacoochee River - Reach 7
30%
25%
20%
15%
Habitat Gain / Loss
10%
10% Flow Reduction
20% Flow Reduction
5%
30% Flow Reduction
40% Flow Reduction
0%
-5%
-10%
-15%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
526
Cricotopus bicinctus 1970-1999
Northern Withlacoochee River - Reach 7
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
527
Tanytarsus sp. C 1940-1969
Northern Withlacoochee River - Reach 7
60%
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
April
August
May
July
February
January
November
March
June
October
September
December
528
Tanytarsus sp. C 1970-1999
Northern Withlacoochee River - Reach 7
100%
80%
60%
Habitat Gain / Loss
20%
0%
-20%
April
August
May
July
February
January
November
March
June
October
September
529 December
Parakiefferiella sp. 1940-1969
Northern Withlacoochee River - Reach 7
30%
20%
10%
Habitat Gain / Loss
-10%
-20%
-30%
April
August
May
July
February
January
November
March
June
October
September
December
530
Parakiefferiella sp. 1970-1999
Northern Withlacoochee River - Reach 7
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
531
Pseudochironomus sp. 1940-1969
Northern Withlacoochee River - Reach 7
40%
30%
20%
10%
Habitat Gain / Loss
0%
10% Flow Reduction
20% Flow Reduction
-10%
30% Flow Reduction
40% Flow Reduction
-20%
-30%
-40%
-50%
-60%
April
August
May
July
February
January
November
March
June
October
September
December
532
Pseudochironomus sp. 1970-1999
Northern Withlacoochee River - Reach 7
80%
60%
40%
Habitat Gain / Loss
-20%
-40%
-60%
April
August
May
July
February
January
November
March
June
October
September
533 December
Stempellina sp. A 1940-1969
Northern Withlacoochee River - Reach 7
250%
200%
Habitat Gain / Loss
150%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
100%
50%
0%
April
August
May
July
February
January
November
March
June
October
September
December
534
Stempellina sp. A 1970-1999
Northern Withlacoochee River - Reach 7
250%
200%
Habitat Gain / Loss
150%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
100%
50%
0%
April
August
May
July
February
January
November
March
June
October
September
535 December
Tvetenia vitracies 1940-1969
Northern Withlacoochee River - Reach 7
120%
100%
80%
60%
Habitat Gain / Loss
0%
-20%
-40%
April
August
May
July
February
January
November
March
June
October
September
December
536
Tvetenia vitracies 1970-1999
Northern Withlacoochee River - Reach 7
200%
150%
Habitat Gain / Loss
100%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
50%
0%
-50%
April
August
May
July
February
January
November
March
June
October
September
537 December
APPENDIX 6. Results of PHABSIM analysis (weighted usable areas) for targeted fish species.
538
Largemouth bass
Northern Withlacoochee River - Reach 1
10000
9000
8000
7000
WUA (sq ft / 1000 linear ft)
6000
Adults
5000
Juveniles
4000
3000
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
539
WUA (sq ft / 1000 linear ft)
0
5000
10000
15000
20000
25000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
540
380
Discharge (cfs)
400
Suwannee bass
450
500
550
600
Northern Withlcoochee River - Reach 1
700
800
900
1000
1100
1200
Adult
Juvenile
WUA (sq ft / 1000 linear ft)
0
5000
10000
15000
20000
25000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
541
380
Discharge (cfs)
Spotted sucker
400
450
500
550
Northern Withlacoochee River - Reach 1
600
700
800
900
1000
1100
1200
Adult
Juvenile
WUA (sq ft / 1000 linear ft)
0
2000
4000
6000
8000
10000
12000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
542
450
500
550
American eel
600
700
800
900
Northern Withlacoochee River - Reach 1
1000
1100
1200
Adult
Largemouth bass
Northern Withlacoochee River - Reach 2
4500
4000
3500
3000
WUA (sq ft / 1000 linear ft)
2500
Adults
2000 Juveniles
1500
1000
500
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
543
Suwannee bass
Northern Withlcoochee River - Reach 2
16000
14000
12000
WUA (sq ft / 1000 linear ft)
10000
Adult
8000
Juvenile
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
544
Spotted sucker
Northern Withlacoochee River - Reach 2
12000
10000
8000
WUA (sq ft / 1000 linear ft)
Adult
6000
Juvenile
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
545
American eel
Northern Withlacoochee River - Reach 2
10000
9000
8000
7000
WUA (sq ft / 1000 linear ft)
6000
5000
Adult
4000
3000
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
546
Largemouth bass
Northern Withlacoochee River - Reach Run 2
16000
14000
12000
WUA (sq ft / 1000 linear ft)
10000
Adults
8000
Juveniles
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
547
Suwannee bass
Northern Withlcoochee River - Reach Run 2
30000
25000
20000
WUA (sq ft / 1000 linear ft)
Adult
15000
Juvenile
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
548
Spotted sucker
Northern Withlacoochee River - Reach Run 2
50000
45000
40000
35000
WUA (sq ft / 1000 linear ft)
30000
Adult
25000
Juvenile
20000
15000
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
549
WUA (sq ft / 1000 linear ft)
0
1000
2000
3000
4000
5000
6000
135 7000
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
550
450
500
550
American eel
600
700
800
900
Northern Withlacoochee River - Reach Run 2
1000
1100
1200
Adult
Largemouth bass
Northern Withlacoochee River - Reach 3
9000
8000
7000
6000
WUA (sq ft / 1000 linear ft)
5000
Adults
4000 Juveniles
3000
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
551
Suwannee bass
Northern Withlcoochee River - Reach 3
14000
12000
10000
WUA (sq ft / 1000 linear ft)
8000
Adult
Juvenile
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
552
Spotted sucker
Northern Withlacoochee River - Reach 3
35000
30000
25000
WUA (sq ft / 1000 linear ft)
20000
Adult
15000 Juvenile
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
553
WUA (sq ft / 1000 linear ft)
0
500
1000
1500
2000
2500
3000
135 3500
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
554
American eel
450
500
550
600
Northern Withlacoochee River - Reach 3
700
800
900
1000
1100
1200
Adult
Largemouth bass
Northern Withlacoochee River - Reach 3.5
12000
10000
8000
WUA (sq ft / 1000 linear ft)
Adults
6000
Juveniles
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
555
Suwannee bass
Northern Withlcoochee River - Reach 3.5
16000
14000
12000
WUA (sq ft / 1000 linear ft)
10000
Adult
8000
Juvenile
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
556
Spotted sucker
Northern Withlacoochee River - Reach 3.5
35000
30000
25000
WUA (sq ft / 1000 linear ft)
20000
Adult
15000 Juvenile
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
557
WUA (sq ft / 1000 linear ft)
0
1000
2000
3000
4000
5000
6000
135 7000
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
558
450
American eel
500
550
600
700
800
Northern Withlacoochee River - Reach 3.5
900
1000
1100
1200
Adult
Largemouth bass
Northern Withlacoochee River - Reach 4
8000
7000
6000
WUA (sq ft / 1000 linear ft)
5000
Adults
4000
Juveniles
3000
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
559
Suwannee bass
Northern Withlcoochee River - Reach 4
20000
18000
16000
14000
WUA (sq ft / 1000 linear ft)
12000
Adult
10000
Juvenile
8000
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
560
WUA (sq ft / 1000 linear ft)
0
5000
10000
15000
20000
25000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
561
380
Discharge (cfs)
400
Spotted sucker
450
500
550
Northern Withlacoochee River - Reach 4
600
700
800
900
1000
1100
1200
Adult
Juvenile
American eel
Northern Withlacoochee River - Reach 4
45000
40000
35000
30000
WUA (sq ft / 1000 linear ft)
25000
Adult
20000
15000
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
562
Largemouth bass
Northern Withlacoochee River - Reach 5
1000
900
800
700
WUA (sq ft / 1000 linear ft)
600
Adults
500
Juveniles
400
300
200
100
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
563
WUA (sq ft / 1000 linear ft)
0
1000
2000
3000
4000
5000
6000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
564
380
Discharge (cfs)
400
450
Suwannee bass
500
550
600
Northern Withlcoochee River - Reach 5
700
800
900
1000
1100
1200
Adult
Juvenile
WUA (sq ft / 1000 linear ft)
0
1000
2000
3000
4000
5000
6000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
565
Discharge (cfs)
400
Spotted sucker
450
500
550
Northern Withlacoochee River - Reach 5
600
700
800
900
1000
1100
1200
Adult
Juvenile
WUA (sq ft / 1000 linear ft)
0
2000
4000
6000
8000
10000
12000
14000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
American eel
566
450
500
550
600
Northern Withlacoochee River - Reach 5
700
800
900
1000
1100
1200
Adult
Largemouth bass
Northern Withlacoochee River - Reach 6
8000
7000
6000
WUA (sq ft / 1000 linear ft)
5000
Adults
4000
Juveniles
3000
2000
1000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
567
WUA (sq ft / 1000 linear ft)
0
2000
4000
6000
8000
10000
12000
135
145
155
165
180
200
220
240
260
280
300
320
340
360
568
380
Discharge (cfs)
400
Suwannee bass
450
500
550
Northern Withlcoochee River - Reach 6
600
700
800
900
1000
1100
1200
Adult
Juvenile
Spotted sucker
Northern Withlacoochee River - Reach 6
20000
18000
16000
14000
WUA (sq ft / 1000 linear ft)
12000
Adult
10000
Juvenile
8000
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
569
American eel
Northern Withlacoochee River - Reach 6
16000
14000
12000
WUA (sq ft / 1000 linear ft)
10000
8000
Adult
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
570
Largemouth bass
Northern Withlacoochee River - Reach Run 7
12000
10000
8000
WUA (sq ft / 1000 linear ft)
Adults
6000
Juveniles
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
571
Suwannee bass
Northern Withlcoochee River - Reach Run 7
20000
18000
16000
14000
WUA (sq ft / 1000 linear ft)
12000
Adult
10000
Juvenile
8000
6000
4000
2000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
572
Spotted sucker
Northern Withlacoochee River - Reach Run 7
35000
30000
25000
WUA (sq ft / 1000 linear ft)
20000
Adult
15000 Juvenile
10000
5000
0
135
145
155
165
180
200
220
240
260
280
300
320
340
360
380
400
450
500
550
600
700
800
900
1000
1100
1200
Discharge (cfs)
573
WUA (sq ft / 1000 linear ft)
0
1000
2000
3000
4000
5000
6000
135 7000
145
155
165
180
200
220
240
260
280
300
320
340
360
380
Discharge (cfs)
400
574
American eel
450
500
550
600
700
Northern Withlacoochee River - Reach Run 7
800
900
1000
1100
1200
Adult
APPENDIX 7. Time series analyses of targeted fish species PHABSIM results.
575
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 1
350%
300%
250%
Habitat Gain / Loss
200%
10% Flow Reduction
20% Flow Reduction
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
December
September
350%
300%
250%
Habitat Gain / Loss
200%
10% Flow Reduction
20% Flow Reduction
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
December
September
576
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 1
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 1
70%
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
577
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 1
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
60% 30% Flow Reduction
40% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 1
140%
120%
100%
Habitat GAin / Loss
80%
10% Flow Reduction
20% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
578
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 1
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 1
40%
30%
20%
Habitat Gain / Loss
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
579
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 1
200%
180%
160%
140%
120%
Habitat Gain/Loss
40%
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 1
200%
180%
160%
140%
120%
Habitat Gain/Loss
60%
40%
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
580
Adult Spotted Sucker (1940-1969)
Withlacoochee River - Shoal 1
60%
50%
40%
30%
Habitat Gain/Loss
20%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
-30%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 1
60%
50%
40%
30%
Habitat Gain/Loss
20%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
0%
-10%
-20%
-30%
-40%
August
July
April
May
June
October
March
January
February
November
September
December
581
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 1
100%
90%
80%
70%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
100%
90%
80%
70%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
582
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
60%
50%
40%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
583
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
Habitat Gain / Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
584
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 2
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 2
80%
70%
60%
50%
Habitat GAin / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
585
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
30%
Habitat Gain / Loss
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
586
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 2
90%
80%
70%
60%
Habitat Gain/Loss
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 2
90%
80%
70%
60%
Habitat Gain/Loss
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
587
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
30%
Habitat Gain/Loss
0%
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 2
60%
50%
40%
30%
Habitat Gain/Loss
0%
-10%
-20%
-30%
August
July
April
May
June
October
March
January
February
November
September
December
588
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 2
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
100%
80%
60%
Habitat Gain/Loss
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
589
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Run 2
70%
60%
50%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
70%
60%
50%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
590
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Run 2
50%
40%
Habitat Gain / Loss
30%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Run 2
50%
40%
Habitat Gain / Loss
30%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
40% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
591
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Run 2
60%
50%
40%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Run 2
60%
50%
40%
Habitat GAin / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
592
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Run 2
50%
40%
30%
Habitat Gain / Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Run 2
50%
40%
30%
Habitat Gain / Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
593
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Run 2
140%
120%
100%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Run 2
140%
120%
100%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
594
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Run 2
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Run 2
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
595
Adult American Eel (1940-1969)
Northern Withlacoochee River - Run 2
40%
30%
Habitat Gain/Loss
10%
0%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
40%
30%
Habitat Gain/Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
596
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 3
60%
50%
40%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
60%
50%
40%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
597
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 3
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 3
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
598
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 3
50%
40%
30%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 3
50%
40%
30%
Habitat GAin / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
599
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 3
30%
20%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 3
30%
20%
Habitat Gain / Loss
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
600
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 3
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 3
80%
70%
60%
50%
Habitat Gain/Loss
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
601
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 3
60%
50%
40%
30%
Habitat Gain/Loss
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 3
60%
50%
40%
30%
Habitat Gain/Loss
0%
-10%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
602
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 3
25%
20%
15%
Habitat Gain/Loss
5%
0%
-5%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
25%
20%
15%
Habitat Gain/Loss
5%
0%
-5%
August
July
April
May
June
October
March
January
February
November
September
December
603
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 3.5
70%
60%
50%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
70%
60%
50%
Habitat Gain / Loss
40%
10% Flow Reduction
20% Flow Reduction
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
604
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 3.5
60%
50%
40%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 3.5
60%
50%
40%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
605
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 3.5
40%
35%
30%
25%
Habitat Gain / Loss
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 3.5
40%
35%
30%
25%
Habitat GAin / Loss
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
December
September
606
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 3.5
40%
35%
30%
25%
Habitat Gain / Loss
20%
10% Flow Reduction
20% Reduction
15%
30% Flow Reduction
40% Flow Reduction
10%
5%
0%
-5%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 3.5
40%
35%
30%
25%
Habitat Gain / Loss
20%
10% Flow Reduction
20% Flow Reduction
15%
30% Flow Reduction
40% Flow Reduction
10%
5%
0%
-5%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
607
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 3.5
300%
250%
200%
Habitat Gain/Loss
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 3.5
300%
250%
200%
Habitat Gain/Loss
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
September
December
608
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 3.5
60%
50%
40%
Habitat Gain/Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 3.5
60%
50%
40%
Habitat Gain/Loss
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
609
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 3.5
35%
30%
25%
20%
Habitat Gain/Loss
15%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
5%
0%
-5%
-10%
-15%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
35%
30%
25%
20%
Habitat Gain/Loss
15%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
5%
0%
-5%
-10%
-15%
August
July
April
May
June
October
March
January
February
November
September
December
610
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 4
450%
400%
350%
300%
Habitat Gain / Loss
250%
10% Flow Reduction
20% Flow Reduction
200% 30% Flow Reduction
40% Flow Reduction
150%
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
December
September
450%
400%
350%
300%
Habitat Gain / Loss
250%
10% Flow Reduction
20% Flow Reduction
200% 30% Flow Reduction
40% Flow Reduction
150%
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
December
September
611
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 4
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 4
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
612
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 4
350%
300%
250%
Habitat Gain / Loss
200%
10% Flow Reduction
20% Flow Reduction
150% 30% Flow Reduction
40% Flow Reduction
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 4
350%
300%
250%
Habitat GAin / Loss
200%
10% Flow Reduction
20% Flow Reduction
100%
50%
0%
August
July
April
May
June
October
March
January
February
November
December
September
613
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 4
60%
50%
40%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 4
60%
50%
40%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
614
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 4
120%
100%
80%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 4
120%
100%
80%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
615
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 4
45%
40%
35%
30%
Habitat Gain/Loss
15%
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 4
45%
40%
35%
30%
Habitat Gain/Loss
15%
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
616
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 4
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
80%
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
617
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 5
120%
100%
80%
Habitat Gain / Loss
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
120%
100%
80%
Habitat Gain / Loss
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
618
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 5
110%
100%
90%
80%
70%
Habitat Gain / Loss
40%
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 5
110%
100%
90%
80%
70%
Habitat Gain / Loss
40%
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
619
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 5
90%
80%
70%
60%
Habitat Gain / Loss
50%
10% Flow Reduction
20% Flow Reduction
40% 30% Flow Reduction
40% Flow Reduction
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 5
90%
80%
70%
60%
Habitat GAin / Loss
50%
10% Flow Reduction
20% Flow Reduction
40% 30% Flow Reduction
40% Flow Reduction
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
December
September
620
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 5
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 5
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
621
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 5
300%
260%
220%
Habitat Gain/Loss
100%
60%
20%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 5
300%
260%
220%
Habitat Gain/Loss
100%
60%
20%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
622
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 5
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 5
70%
60%
50%
Habitat Gain/Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
623
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 5
180%
160%
140%
120%
Habitat Gain/Loss
100%
10% Flow Reduction
20% Flow Reduction
80%
30% Flow Reduction
40% Flow Reduction
60%
40%
20%
0%
-20%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
180%
160%
140%
120%
Habitat Gain/Loss
100%
10% Flow Reduction
20% Flow Reduction
80%
30% Flow Reduction
40% Flow Reduction
60%
40%
20%
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
624
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 6
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
625
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Shoal 6
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Shoal 6
80%
70%
60%
50%
Habitat Gain / Loss
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
626
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 6
30%
25%
20%
15%
Habitat Gain / Loss
0%
-5%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Shoal 6
30%
25%
20%
15%
Habitat GAin / Loss
0%
-5%
-10%
August
July
April
May
June
October
March
January
February
November
December
September
627
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Shoal 6
30%
25%
20%
Habitat Gain / Loss
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Shoal 6
30%
25%
20%
Habitat Gain / Loss
10%
5%
0%
August
July
April
May
June
October
March
January
February
November
September
December
628
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 6
140%
120%
100%
Habitat Gain/Loss
80%
10% Flow Reduction
20% Flow Reduction
30% Flow Reduction
60%
40% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 6
140%
120%
100%
Habitat Gain/Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
629
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Shoal 6
35%
30%
25%
20%
Habitat Gain/Loss
5%
0%
-5%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Shoal 6
35%
30%
25%
20%
Habitat Gain/Loss
5%
0%
-5%
August
July
April
May
June
October
March
January
February
November
September
December
630
Adult American Eel (1940-1969)
Northern Withlacoochee River - Shoal 6
20%
18%
16%
14%
Habitat Gain/Loss
12%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
8%
6%
4%
2%
0%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
20%
18%
16%
14%
Habitat Gain/Loss
12%
10% Flow Reduction
20% Flow Reduction
10%
30% Flow Reduction
40% Flow Reduction
8%
6%
4%
2%
0%
August
July
April
May
June
October
March
January
February
November
September
December
631
Juvnile Largemouth Bass (1940-1969)
Northern Withlacoochee River - Run 7
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
140%
120%
100%
Habitat Gain / Loss
80%
10% Flow Reduction
20% Flow Reduction
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
December
September
632
Adult Largemouth Bass (1940-1969)
Northern Withlacoochee River - Run 7
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Largemouth Bass (1970-1999)
Northern Withlacoochee River - Run 7
120%
100%
80%
Habitat Gain / Loss
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
633
Juvenile Suwannee Bass (1940-1969)
Northern Withlacoochee River - Run 7
100%
80%
60%
Habitat Gain / Loss
0%
-20%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Suwannee Bass 1970-1999
Northern Withlacoochee River - Run 7
100%
80%
60%
Habitat GAin / Loss
0%
-20%
August
July
April
May
June
October
March
January
February
November
December
September
634
Adult Suwannee Bass (1940-1969)
Northern Withlacoochee River - Run 7
100%
90%
80%
70%
60%
Habitat Gain / Loss
30%
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Suwannee Bass (1970-1999)
Northern Withlacoochee River - Run 7
100%
90%
80%
70%
60%
Habitat Gain / Loss
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
635
Juvenile Spotted Sucker (1940-1969)
Northern Withlacoochee River - Run 7
180%
160%
140%
120%
Habitat Gain/Loss
60%
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
Juvenile Spotted Sucker (1970-1999)
Northern Withlacoochee River - Run 7
180%
160%
140%
120%
Habitat Gain/Loss
60%
40%
20%
0%
August
July
April
May
June
October
March
January
February
November
September
December
636
Adult Spotted Sucker (1940-1969)
Northern Withlacoochee River - Run 7
110%
100%
90%
80%
70%
Habitat Gain/Loss
30%
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
Adult Spotted Sucker (1970-1999)
Northern Withlacoochee River - Run 7
110%
100%
90%
80%
70%
Habitat Gain/Loss
30%
20%
10%
0%
-10%
August
July
April
May
June
October
March
January
February
November
September
December
637
Adult American Eel (1940-1969)
Northern Withlacoochee River - Run 7
100%
90%
80%
70%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
August
July
October
April
May
June
Janua ry
November
March
February
December
September
100%
90%
80%
70%
Habitat Gain/Loss
60%
10% Flow Reduction
20% Flow Reduction
50%
30% Flow Reduction
40% Flow Reduction
40%
30%
20%
10%
0%
August
July
April
May
June
October
March
January
February
November
September
December
638