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Fermentative Metabolism
NADH NAD Fructose-6-P
1
Citric Acid Cycle oxidation of malate
NAD+
Malate + NAD+ Oxaloacetate + NADH
Malate
dehydrogenase
Start Mon
NAD+
Malate + NAD+ Pyruvate + CO2 + NADH
malic enzyme
Functions
1. Complete oxidation of Krebs cycle intermediates
Electron &
Proton
carrier
FMN &
Fe-S
proteins
e-
Fe - S
2e- Fe-S
proteins
proteins
b&c
cytochromes
e- e-
Cu centers
cytochromes
a1 & a3
3. Response to stress
Draining Ubiquinone pool prevents
Voodoo lily superoxide/hydroxide buildup
Skunk cabbage
2
Lipid Storage
Lipid Functions 1. Oil bodies store lipids = spherosomes or oleosomes
Half-bilayer membrane
1. Carbon storage single layer of phospholipids formed from ER
Seeds e.g. commercial oils: sunflower, soybean, peanut, cotton formed within ER membrane bilayer
Fruits e.g. avocado, olives Oleosin proteins stabilize membrane
2. Energy Storage same as above
3. Energy harvest chlorophylls & carotenoids
4. Membranes phospholipids
5. Protection waxes
6. Hormones isopreness precursors to some hormones (ABA & GA)
metabolic products (natural rubber)
sterols & tocopherols Oil Body
7. Intracellular signalling
linolenic acid jasmonate activates defenses against
fungal & insect pathogens
anther & pollen development
Triacylglycerols
1. Glycerol + 3 fatty acids (acyl groups)
Fatty acids: 12 20 carbons, mostly 16-18
Oils unsaturated fatty acids
Fats mostly saturated fatty acids
Glycerophospholipids
3
Lipid Conversion to Carbohydrates
Biosynthesis of Glycerophospholipids & Glyceroglycolipids
-oxidation of fatty acids
Acetyl CoA Oil Triacylglycerol hydrolysis to free fatty acids
Acyl Carrier Protein 100% in Bodies
Condensation to fatty acids-ACP Chloroplast
Oilseed
Glycerol-3-P
Sucrose
Growth
Glycerophospholipids
Different Glucose Mitochondrion
Acyl
Carriers
Gluconeogenesis
Malate Oxaloacetate
Cytosol
Glyceroglycopholipids
END
Respiration &
Lipid Metabolism