_______________________________________________________________

QUADRAT DIVISION 1 Number of quadrates in cluster = 48

eigenvalue = 0.5501 number of iterations = 2
Indicators and their sign
Aga [-];
The maximum indicator score for the negative group = -1
The minimum indicator score for the positive group = 0
Negative group: 2 Number of objects = 24 comprising:
1, 1, 1, 1, 1, 1, 1, 1, 2, 2, 2, 2, 2, 2, 2, 2, 3, 3, 3, 3, 3, 3, 3, 3,
The borderline negative group: Number of objects = 1 comprising:
3,
The positive group: 3 Number of objects = 24 comprising:
4, 4, 4, 4, 4, 4, 4, 4, 5, 5, 5, 5, 5, 5, 5, 5, 6, 6, 6, 6, 6, 6, 6, 6,
Species preferring the negative group of quadrats
Chim 1 (22, 4) Gen 1 (10, 0) Aga 1 (24, 0) Diple 1 (21, 2) Ortho 1 (11, 0) Cae 1
(15, 0) Lepi 1 (20, 0) Anis 1 (5, 0) Chim 2 (22, 4) Gen 2 (10, 0) Aga 2 (24, 0)
Diple 2 (21, 2) Ortho 2 (11, 0) Cae 2 (15, 0) Lepi 2 (20, 0) Anis 2 (5, 0) Chim
3 (10, 1) Aga 3 (20, 0) Diple 3 (9, 0) Ortho 3 (5, 0) Cae 3 (11, 0) Lepi 3 (16,
0) Chim 4 (9, 0) Aga 4 (16, 0) Cae 4 (7, 0) Lepi 4 (11, 0) Aga 5 (12, 0)
Species biased towards the positive group of quadrats
Genus 1 1 (0, 12) Tino 1 (7, 19) Seto 1 (0, 10) Genus 1 2 (0, 12) Tino 2 (7, 19)
Seto 2 (0, 10) Genus 1 3 (0, 6) Tino 3 (6, 16) Seto 3 (0, 6) Tino 4 (5, 12) Tin
o 5 (2, 7)
Species with no quadrat preference
Cheu 1 (24, 24) Cheu 2 (24, 24) Cheu 3 (23, 24) Cheu 4 (23, 24) Cheu 5 (20, 22)
END OF LEVEL 1
TWINSPAN QUADRAT CLASSIFICATION LEVEL 1
_______________________________________________________________
QUADRAT DIVISION 2 Number of quadrates in cluster = 24
eigenvalue = 0.275 number of iterations = 3
Indicators and their sign
Tino [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 4 Number of objects = 17 comprising:
1, 1, 1, 1, 1, 1, 1, 1, 2, 2, 2, 2, 2, 2, 2, 2, 3,
The borderline negative group: Number of objects = 2 comprising:
2, 3,
The positive group: 5 Number of objects = 7 comprising:
3, 3, 3, 3, 3, 3, 3,

Species preferring the negative group of quadrats

Gen 1 (9, 1) Ortho 1 (11, 0) Helico 1 (4, 0) Anti 1 (4, 0) Cae 1 (15, 0) Lepi 1
(17, 3) Anis 1 (5, 0) Gen 2 (9, 1) Ortho 2 (11, 0) Helico 2 (4, 0) Anti 2 (4, 0)
Cae 2 (15, 0) Lepi 2 (17, 3) Anis 2 (5, 0) Chim 3 (9, 1) Ortho 3 (5, 0) Cae 3 (
11, 0) Lepi 3 (15, 1) Chim 4 (8, 1) Cae 4 (7, 0) Lepi 4 (10, 1) Chim 5 (4, 0)
Species biased towards the positive group of quadrats
Tino 1 (0, 7) Tino 2 (0, 7) Tino 3 (0, 6) Tino 4 (0, 5) Aga 5 (6, 6) Tino 5 (0,
2)
Species with no quadrat preference
Chim 1 (15, 7) Aga 1 (17, 7) Diple 1 (15, 6) Cheu 1 (17, 7) Chim 2 (15, 7) Aga 2
(17, 7) Diple 2 (15, 6) Cheu 2 (17, 7) Aga 3 (13, 7) Diple 3 (7, 2) Cheu 3 (16,
7) Aga 4 (9, 7) Cheu 4 (16, 7) Cheu 5 (14, 6)
_______________________________________________________________
QUADRAT DIVISION 3 Number of quadrates in cluster = 24
eigenvalue = 0.1908 number of iterations = 6
Indicators and their sign
Seto [+];
Genus 1 [-];
Tino [+];
Diple [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 6 Number of objects = 13 comprising:
4, 4, 4, 5, 5, 5, 5, 5, 5, 6, 6, 6, 6,
The borderline negative group: Number of objects = 1 comprising:
5,
The positive group: 7 Number of objects = 11 comprising:
4, 4, 4, 4, 4, 5, 5, 6, 6, 6, 6,
The borderline positive group: Number of objects = 3 comprising:
4, 6, 6,
The misclassified positive:Number of objects = 1 comprising:
5,
Species preferring the negative group of quadrats
Chim 1 (3, 1) Genus 1 1 (10, 2) Chim 2 (3, 1) Genus 1 2 (10, 2) Genus 1 3 (6, 0)
Species biased towards the positive group of quadrats
Seto 1 (2, 8) Seto 2 (2, 8) Seto 3 (1, 5) Tino 4 (3, 9) Seto 4 (1, 3) Tino 5 (2,
5)
Species with no quadrat preference

Cheu 1 (13, 11) Tino 1 (9, 10) Cheu 2 (13, 11) Tino 2 (9, 10) Cheu 3 (13, 11) Ti
no 3 (6, 10) Cheu 4 (13, 11) Cheu 5 (12, 10)
END OF LEVEL 2
TWINSPAN QUADRAT CLASSIFICATION LEVEL 2
_______________________________________________________________
QUADRAT DIVISION 4 Number of quadrates in cluster = 17
eigenvalue = 0.1817 number of iterations = 4
Indicators and their sign
Anis [-];
Gen [-];
Cae [-];
The maximum indicator score for the negative group = -2
The minimum indicator score for the positive group = -1
Negative group: 8 Number of objects = 6 comprising:
1, 1, 1, 2, 2, 2,
The positive group: 9 Number of objects = 11 comprising:
1, 1, 1, 1, 1, 2, 2, 2, 2, 2, 3,
The borderline positive group: Number of objects = 1 comprising:
1,
Species preferring the negative group of quadrats
Gen 1 (6, 3) Apsil 1 (3, 0) Ulme 1 (2, 0) Anis 1 (5, 0) Gen 2 (6, 3) Apsil 2 (3,
0) Ulme 2 (2, 0) Anis 2 (5, 0) Diple 3 (4, 3) Diple 4 (2, 0) Cae 4 (5, 2) Chim
5 (3, 1) Cae 5 (3, 0)
Species biased towards the positive group of quadrats
Ortho 1 (2, 9) Helico 1 (0, 4) Anti 1 (0, 4) Ortho 2 (2, 9) Helico 2 (0, 4) Anti
2 (0, 4) Ortho 3 (0, 5) Ortho 4 (0, 3)
Species with no quadrat preference
Chim 1 (6, 9)
6, 11) Chim 2
epi 2 (6, 11)
9) Chim 4 (4,
5, 9)

Aga 1 (6, 11) Diple 1 (6, 9) Cheu 1 (6, 11) Cae 1 (5, 10) Lepi 1 (
(6, 9) Aga 2 (6, 11) Diple 2 (6, 9) Cheu 2 (6, 11) Cae 2 (5, 10) L
Chim 3 (4, 5) Aga 3 (5, 8) Cheu 3 (6, 10) Cae 3 (5, 6) Lepi 3 (6,
4) Aga 4 (4, 5) Cheu 4 (6, 10) Lepi 4 (5, 5) Aga 5 (2, 4) Cheu 5 (

_______________________________________________________________
QUADRAT DIVISION 5 Number of quadrates in cluster = 7
eigenvalue = 0.1596 number of iterations = 3
Indicators and their sign
Diple [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 10 Number of objects = 1 comprising:
3,
The positive group: 11 Number of objects = 6 comprising:

3, 3, 3, 3, 3, 3,
Species preferring the negative group of quadrats
Lepi 1 (1, 2) Lepi 2 (1, 2) Chim 3 (1, 0) Lepi 3 (1, 0) Chim 4 (1, 0) Lepi 4 (1,
0)
Species biased towards the positive group of quadrats
Diple 1 (0, 6) Diple 2 (0, 6) Diple 3 (0, 2) Tino 5 (0, 2)
Species with no quadrat preference
Chim 1 (1, 6) Aga 1 (1, 6) Cheu 1 (1, 6) Tino 1 (1, 6) Chim 2 (1, 6) Aga 2 (1, 6
) Cheu 2 (1, 6) Tino 2 (1, 6) Aga 3 (1, 6) Cheu 3 (1, 6) Tino 3 (1, 5) Aga 4 (1,
6) Cheu 4 (1, 6) Tino 4 (1, 4) Aga 5 (1, 5) Cheu 5 (1, 5)
_______________________________________________________________
QUADRAT DIVISION 6 Number of quadrates in cluster = 13
eigenvalue = 0.2355 number of iterations = 2
Indicators and their sign
Tino [-];
The maximum indicator score for the negative group = -1
The minimum indicator score for the positive group = 0
Negative group: 12 Number of objects = 6 comprising:
4, 4, 4, 5, 5, 5,
The positive group: 13 Number of objects = 7 comprising:
5, 5, 5, 6, 6, 6, 6,
The borderline positive group: Number of objects = 1 comprising:
5,
Species preferring the negative group of quadrats
Chim 1 (3, 0) Tino 1 (6, 3) Chim 2 (3, 0) Tino 2 (6, 3) Tino 3 (6, 0) Tino 4 (3,
0) Tino 5 (2, 0)
Species biased towards the positive group of quadrats
Seto 1 (0, 2) Seto 2 (0, 2)
Species with no quadrat preference
Genus 1 1 (5, 5) Cheu 1 (6, 7) Genus 1 2 (5, 5) Cheu 2 (6, 7) Genus 1 3 (2, 4) C
heu 3 (6, 7) Cheu 4 (6, 7) Cheu 5 (5, 7)
_______________________________________________________________
QUADRAT DIVISION 7 Number of quadrates in cluster = 11
eigenvalue = 0.2529 number of iterations = 3
Indicators and their sign
Seto [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 14 Number of objects = 8 comprising:
4, 4, 4, 4, 5, 6, 6, 6,

The borderline negative group: Number of objects = 1 comprising:

6,
The positive group: 15 Number of objects = 3 comprising:
4, 5, 6,
Species preferring the negative group of quadrats
Diple 1 (2, 0) Diple 2 (2, 0)
Species biased towards the positive group of quadrats
Chim 1 (0, 1) Genus 1 1 (0, 2) Chim 2 (0, 1) Genus 1 2 (0, 2) Chim 3 (0, 1) Seto
3 (2, 3) Seto 4 (0, 3) Seto 5 (0, 2)
Species with no quadrat preference
Cheu 1 (8, 3) Tino 1 (8, 2) Seto 1 (5, 3) Cheu 2 (8, 3) Tino 2 (8, 2) Seto 2 (5,
3) Cheu 3 (8, 3) Tino 3 (8, 2) Cheu 4 (8, 3) Tino 4 (7, 2) Cheu 5 (7, 3) Tino 5
(4, 1)
END OF LEVEL 3
TWINSPAN QUADRAT CLASSIFICATION LEVEL 3
_______________________________________________________________
QUADRAT DIVISION 8 Number of quadrates in cluster = 6
eigenvalue = 0.145 number of iterations = 3
Indicators and their sign
Ulme [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 16 Number of objects = 4 comprising:
1, 1, 1, 2,
The positive group: 17 Number of objects = 2 comprising:
2, 2,
Species preferring the negative group of quadrats
Apsil 1 (3, 0) Cae 1 (4, 1) Apsil 2 (3, 0) Cae 2 (4, 1) Chim 3 (4, 0) Cae 3 (4,
1) Chim 4 (4, 0) Diple 4 (2, 0) Cae 4 (4, 1) Chim 5 (3, 0) Cae 5 (3, 0) Lepi 5 (
1, 0)
Species biased towards the positive group of quadrats
Ortho 1 (1, 1) Ulme 1 (0, 2) Ortho 2 (1, 1) Ulme 2 (0, 2) Gen 3 (0, 1) Diple 3 (
2, 2) Aga 4 (2, 2) Aga 5 (1, 1)
Species with no quadrat preference
Chim 1 (4, 2) Gen 1 (4, 2) Aga 1 (4, 2) Diple 1 (4, 2) Cheu 1 (4, 2) Lepi 1 (4,
2) Anis 1 (3, 2) Chim 2 (4, 2) Gen 2 (4, 2) Aga 2 (4, 2) Diple 2 (4, 2) Cheu 2 (
4, 2) Lepi 2 (4, 2) Anis 2 (3, 2) Aga 3 (3, 2) Cheu 3 (4, 2) Lepi 3 (4, 2) Cheu
4 (4, 2) Lepi 4 (3, 2) Cheu 5 (3, 2)

_______________________________________________________________
QUADRAT DIVISION 9 Number of quadrates in cluster = 11
eigenvalue = 0.1829 number of iterations = 3
Indicators and their sign
Gen [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 18 Number of objects = 8 comprising:
1, 1, 1, 2, 2, 2, 2, 2,
The borderline negative group: Number of objects = 1 comprising:
2,
The positive group: 19 Number of objects = 3 comprising:
1, 1, 3,
Species preferring the negative group of quadrats
Helico 1 (4, 0) Helico 2 (4, 0) Diple 3 (3, 0) Cae 4 (2, 0) Lepi 5 (2, 0)
Species biased towards the positive group of quadrats
Gen 1 (0, 3) Anti 1 (2, 2) Gen 2 (0, 3) Anti 2 (2, 2) Gen 3 (0, 1) Anti 3 (0, 1)
Chim 4 (2, 2) Gen 4 (0, 1) Aga 4 (2, 3) Ortho 4 (1, 2) Chim 5 (0, 1) Aga 5 (1,
3)
Species with no quadrat preference
Chim 1 (7, 2) Aga 1 (8, 3) Diple 1 (7, 2) Cheu 1 (8, 3) Ortho 1 (7, 2) Cae 1 (8,
2) Lepi 1 (8, 3) Chim 2 (7, 2) Aga 2 (8, 3) Diple 2 (7, 2) Cheu 2 (8, 3) Ortho
2 (7, 2) Cae 2 (8, 2) Lepi 2 (8, 3) Chim 3 (3, 2) Aga 3 (5, 3) Cheu 3 (7, 3) Ort
ho 3 (3, 2) Cae 3 (5, 1) Lepi 3 (7, 2) Cheu 4 (7, 3) Lepi 4 (4, 1) Cheu 5 (7, 2)
_______________________________________________________________
QUADRAT DIVISION 10 Number of quadrates in cluster = 1
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 11 Number of quadrates in cluster = 6
eigenvalue = 0.1095 number of iterations = 2
Indicators and their sign
Lepi [-];
The maximum indicator score for the negative group = -1
The minimum indicator score for the positive group = 0
Negative group: 22 Number of objects = 2 comprising:
3, 3,
The positive group: 23 Number of objects = 4 comprising:
3, 3, 3, 3,
The borderline positive group: Number of objects = 2 comprising:
3, 3,

Species preferring the negative group of quadrats

Lepi 1 (2, 0) Lepi 2 (2, 0) Diple 3 (1, 1) Tino 5 (1, 1)
Species biased towards the positive group of quadrats
Gen 1 (0, 1) Gen 2 (0, 1) Tino 3 (1, 4) Cheu 5 (1, 4)
Species with no quadrat preference
Chim 1 (2, 4) Aga 1 (2, 4) Diple 1 (2, 4) Cheu 1 (2, 4) Tino 1 (2, 4) Chim 2 (2,
4) Aga 2 (2, 4) Diple 2 (2, 4) Cheu 2 (2, 4) Tino 2 (2, 4) Aga 3 (2, 4) Cheu 3
(2, 4) Aga 4 (2, 4) Cheu 4 (2, 4) Tino 4 (1, 3) Aga 5 (2, 3)
_______________________________________________________________
QUADRAT DIVISION 12 Number of quadrates in cluster = 6
eigenvalue = 0.116 number of iterations = 2
Indicators and their sign
Tino [-];
The maximum indicator score for the negative group = -1
The minimum indicator score for the positive group = 0
Negative group: 24 Number of objects = 2 comprising:
4, 4,
The positive group: 25 Number of objects = 4 comprising:
4, 5, 5, 5,
The borderline positive group: Number of objects = 2 comprising:
4, 5,
Species preferring the negative group of quadrats
Chim 1 (2, 1) Chim 2 (2, 1) Genus 1 3 (1, 1) Tino 4 (2, 1) Tino 5 (2, 0)
Species biased towards the positive group of quadrats

Species with no quadrat preference

Genus 1 1 (2, 3) Cheu 1 (2, 4) Tino 1 (2, 4) Genus 1 2 (2, 3) Cheu 2 (2, 4) Tino
2 (2, 4) Cheu 3 (2, 4) Tino 3 (2, 4) Cheu 4 (2, 4) Cheu 5 (2, 3)
_______________________________________________________________
QUADRAT DIVISION 13 Number of quadrates in cluster = 7
eigenvalue = 0.2569 number of iterations = 2
Indicators and their sign
Tino [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 26 Number of objects = 4 comprising:
6, 6, 6, 6,
The positive group: 27 Number of objects = 3 comprising:

5, 5, 5,
Species preferring the negative group of quadrats

Species biased towards the positive group of quadrats

Genus 1 1 (2, 3) Tino 1 (0, 3) Seto 1 (0, 2) Genus 1 2 (2, 3) Tino 2 (0, 3) Seto
2 (0, 2) Genus 1 3 (1, 3) Seto 3 (0, 1) Genus 1 4 (0, 1) Seto 4 (0, 1) Seto 5 (
0, 1)
Species with no quadrat preference
Cheu 1 (4, 3) Cheu 2 (4, 3) Cheu 3 (4, 3) Cheu 4 (4, 3) Cheu 5 (4, 3)
_______________________________________________________________
QUADRAT DIVISION 14 Number of quadrates in cluster = 8
eigenvalue = 0.1612 number of iterations = 2
Indicators and their sign
Tino [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 28 Number of objects = 1 comprising:
4,
The positive group: 29 Number of objects = 7 comprising:
4, 4, 4, 5, 6, 6, 6,
Species preferring the negative group of quadrats
Diple 1 (1, 1) Diple 2 (1, 1)
Species biased towards the positive group of quadrats
Seto 1 (0, 5) Seto 2 (0, 5) Seto 3 (0, 2) Tino 4 (0, 7) Cheu 5 (0, 7) Tino 5 (0,
4)
Species with no quadrat preference
Cheu 1 (1, 7) Tino 1 (1, 7) Cheu 2 (1, 7) Tino 2 (1, 7) Cheu 3 (1, 7) Tino 3 (1,
7) Cheu 4 (1, 7)
_______________________________________________________________
QUADRAT DIVISION 15 Number of quadrates in cluster = 3
No division was undertaken as there are too few objects
_______________________________________________________________
END OF LEVEL 4
TWINSPAN QUADRAT CLASSIFICATION LEVEL 4
_______________________________________________________________
QUADRAT DIVISION 16 Number of quadrates in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 17 Number of quadrates in cluster = 2
No division was undertaken as there are too few objects

_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 18 Number of quadrates in cluster = 8
eigenvalue = 0.1516 number of iterations = 2
Indicators and their sign
Cae [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 36 Number of objects = 6 comprising:
1, 2, 2, 2, 2, 2,
The positive group: 37 Number of objects = 2 comprising:
1, 1,
Species preferring the negative group of quadrats
Diple 1 (6, 1) Anti 1 (2, 0) Diple 2 (6, 1) Anti 2 (2, 0) Cheu 3 (6, 1) Cheu 4 (
6, 1) Cheu 5 (6, 1)
Species biased towards the positive group of quadrats
Helico 1 (2, 2) Helico 2 (2, 2) Aga 3 (3, 2) Ortho 3 (1, 2) Cae 3 (3, 2) Chim 4
(1, 1) Aga 4 (1, 1) Ortho 4 (0, 1) Cae 4 (0, 2) Lepi 4 (2, 2) Aga 5 (0, 1) Lepi
5 (0, 2)
Species with no quadrat preference
Chim 1 (5, 2) Aga 1 (6, 2) Cheu 1 (6, 2) Ortho 1 (5, 2) Cae 1 (6, 2) Lepi 1 (6,
2) Chim 2 (5, 2) Aga 2 (6, 2) Cheu 2 (6, 2) Ortho 2 (5, 2) Cae 2 (6, 2) Lepi 2 (
6, 2) Chim 3 (2, 1) Diple 3 (2, 1) Lepi 3 (5, 2)
_______________________________________________________________
QUADRAT DIVISION 19 Number of quadrates in cluster = 3
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 22 Number of quadrates in cluster = 2
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 23 Number of quadrates in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 24 Number of quadrates in cluster = 2
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 25 Number of quadrates in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 26 Number of quadrates in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 27 Number of quadrates in cluster = 3
No division was undertaken as there are too few objects

_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 28 Number of quadrates in cluster = 1
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 29 Number of quadrates in cluster = 7
eigenvalue = 0.1194 number of iterations = 2
Indicators and their sign
Diple [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 58 Number of objects = 6 comprising:
4, 4, 5, 6, 6, 6,
The positive group: 59 Number of objects = 1 comprising:
4,
Species preferring the negative group of quadrats

Species biased towards the positive group of quadrats

Diple 1 (0, 1) Diple 2 (0, 1) Seto 3 (1, 1) Tino 5 (3, 1)
Species with no quadrat preference
Cheu 1 (6, 1) Tino 1 (6, 1) Seto 1 (4, 1) Cheu 2 (6, 1) Tino 2 (6, 1) Seto 2 (4,
1) Cheu 3 (6, 1) Tino 3 (6, 1) Cheu 4 (6, 1) Tino 4 (6, 1) Cheu 5 (6, 1)
END OF LEVEL 5
TWINSPAN QUADRAT CLASSIFICATION LEVEL 5
_______________________________________________________________
QUADRAT DIVISION 36 Number of quadrates in cluster = 6
eigenvalue = 0.1365 number of iterations = 2
Indicators and their sign
Anti [+];
The maximum indicator score for the negative group = 0
The minimum indicator score for the positive group = 1
Negative group: 72 Number of objects = 4 comprising:
1, 2, 2, 2,
The positive group: 73 Number of objects = 2 comprising:
2, 2,
Species preferring the negative group of quadrats
Chim 1 (4, 1) Chim 2 (4, 1) Cae 3 (3, 0) Lepi 3 (4, 1) Aga 4 (1, 0) Lepi 4 (2, 0
)
Species biased towards the positive group of quadrats
Helico 1 (1, 1) Anti 1 (0, 2) Helico 2 (1, 1) Anti 2 (0, 2) Chim 3 (1, 1) Diple

3 (1, 1) Ortho 3 (0, 1) Chim 4 (0, 1)

Species with no quadrat preference
Aga 1 (4, 2) Diple 1 (4, 2) Cheu 1 (4, 2) Ortho 1 (3, 2) Cae 1 (4, 2) Lepi 1 (4,
2) Aga 2 (4, 2) Diple 2 (4, 2) Cheu 2 (4, 2) Ortho 2 (3, 2) Cae 2 (4, 2) Lepi 2
(4, 2) Aga 3 (2, 1) Cheu 3 (4, 2) Cheu 4 (4, 2) Cheu 5 (4, 2)
_______________________________________________________________
QUADRAT DIVISION 37 Number of quadrates in cluster = 2
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
QUADRAT DIVISION 58 Number of quadrates in cluster = 6
eigenvalue = 0.08536 number of iterations = 1
Indicators and their sign
Tino [-];
The maximum indicator score for the negative group = -1
The minimum indicator score for the positive group = 0
Negative group: 116 Number of objects = 3 comprising:
4, 5, 6,
The positive group: 117 Number of objects = 3 comprising:
4, 6, 6,
The borderline positive group: Number of objects = 1 comprising:
6,
Species preferring the negative group of quadrats
Tino 5 (3, 0)
Species biased towards the positive group of quadrats
Seto 3 (0, 1)
Species with no quadrat preference
Cheu 1 (3, 3) Tino 1 (3, 3) Seto 1 (2, 2) Cheu 2 (3, 3) Tino 2 (3, 3) Seto 2 (2,
2) Cheu 3 (3, 3) Tino 3 (3, 3) Cheu 4 (3, 3) Tino 4 (3, 3) Cheu 5 (3, 3)
_______________________________________________________________
QUADRAT DIVISION 59 Number of quadrates in cluster = 1
No division was undertaken as there are too few objects
_______________________________________________________________
END OF LEVEL 6
THIS IS THE END OF THE DIVISIONS REQUESTED
TWINSPAN QUADRAT CLASSIFICATION LEVEL 6
_______________________________________________________________
SPECIES DIVISION 1 Number of species in cluster = 16
eigenvalue = 0.9277 number of iterations = 2
Negative group: 2 Number of objects = 12 comprising:
Chim, Gen, Aga, Diple, Ortho, Helico, Anti, Apsil, Ulme, Cae, Lepi, Anis,

The positive group: 3 Number of objects = 4 comprising:

Genus 1, Cheu, Tino, Seto,
END OF LEVEL 1
TWINSPAN SPECIES CLASSIFICATION LEVEL 1
_______________________________________________________________
SPECIES DIVISION 2 Number of species in cluster = 12
eigenvalue = 0.1948 number of iterations = 2
Negative group: 4 Number of objects = 9 comprising:
Gen, Ortho, Helico, Anti, Apsil, Ulme, Cae, Lepi, Anis,
The positive group: 5 Number of objects = 3 comprising:
Chim, Aga, Diple,
_______________________________________________________________
SPECIES DIVISION 3 Number of species in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
END OF LEVEL 2
TWINSPAN SPECIES CLASSIFICATION LEVEL 2
_______________________________________________________________
SPECIES DIVISION 4 Number of species in cluster = 9
eigenvalue = 0.08467 number of iterations = 2
Negative group: 8 Number of objects = 4 comprising:
Gen, Apsil, Ulme, Anis,
The positive group: 9 Number of objects = 5 comprising:
Ortho, Helico, Anti, Cae, Lepi,
_______________________________________________________________
SPECIES DIVISION 5 Number of species in cluster = 3
No division was undertaken as there are too few objects
_______________________________________________________________
END OF LEVEL 3
TWINSPAN SPECIES CLASSIFICATION LEVEL 3
_______________________________________________________________
SPECIES DIVISION 8 Number of species in cluster = 4
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
SPECIES DIVISION 9 Number of species in cluster = 5
eigenvalue = 0.0616 number of iterations = 1
Negative group: 18 Number of objects = 2 comprising:
Cae, Lepi,
The positive group: 19 Number of objects = 3 comprising:
Ortho, Helico, Anti,

END OF LEVEL 4
TWINSPAN SPECIES CLASSIFICATION LEVEL 4
_______________________________________________________________
SPECIES DIVISION 18 Number of species in cluster = 2
No division was undertaken as there are too few objects
_______________________________________________________________
_______________________________________________________________
SPECIES DIVISION 19 Number of species in cluster = 3
No division was undertaken as there are too few objects
_______________________________________________________________
THIS IS THE END OF THE DIVISIONS REQUESTED

ORDER OF SPECIES INCLUDING RARER ONES

2, 11, 12, 16, 13, 14, 7, 8, 10, 1, 3, 5, 4, 6, 9, 15,
ORDER OF SAMPLES
4, 6, 8, 16, 14, 15, 2, 9, 11, 13, 10, 12, 1, 3, 5, 7, 21, 22, 20, 24, 17, 18, 1
9, 23, 27, 29, 26, 33, 35, 37, 41, 43, 46, 48, 38, 39, 40, 28, 32, 34, 47, 31, 4
2, 45, 30, 25, 36, 44,
10000000000
Gen
0
0
0
0
10000000000
11
Apsil
0
0
0
0
0
0
0
0
0
10000000000
12
Ulme
0
0
0
0
0
0
0
0
10000000000
16
Anis
0
0
0
0
0
0
0
0
10010000000
13
Cae
9
4
0
0
0
0
0
0
10010000000
14
Lepi
2
0
0
0
0

2
0
0
0
0

2
0
0
0
0

2
0
0
0
0

2
0
2
0
0

2
11
0
0
0

7
4
0
0
0

0
2
0
0
0

0
0
0
0
0

0
0
0
0

2
0
0
0
0

2
0
0
0
0

4
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

2
0
0
0
0

2
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

2
0
0
0
0

2
0
0
0
0

2
0
0
0
0

2
0
0
0
0

2
0
0
0
0

0
0
0
0
0

0
0
0
0
0

11
4
0
0
0

22
2
0
0
0

31
18
0
0
0

22
16
0
0
0

0
2
0
0
0

11
7
0
0
0

7
0
0
0
0

9
0
0
0
0

13

33

13

11

13

11

11
9
4
4
0
0
0
0
10011000000
7
Ortho
2
4
0
0
0
0
0
0
0
10011000000
8
Helico
0
2
0
0
0
0
0
0
0
10011000000
10
Anti
0
0
0
0
0
0
0
0
10100000000
1
Chim
2
2
2
2
0
0
0
0
10100000000
3
Aga
2
2
71
42
0
0
0
0
10100000000
5
Diple
2
4
0
4
0
0
2
0
0
11000000000
4
Genus 1
0
0
0
0
0
4
0
0
0
11000000000
6
Cheu
47
27
13
42
18
238
218
131
11000000000
9
Tino
0
0
4
24

2
0
0
0

7
0
0
0

40
0
0
0

22
0
0
0

11
0
0
0

4
0
0
0

7
0
0
0

16
0
0
0

0
0
0
0

4
4
0
0
0

0
7
0
0
0

0
11
0
0
0

0
9
0
0
0

0
18
0
0
0

4
11
0
0
0

2
0
0
0
0

0
0
0
0

0
4
0
0
0

0
0
0
0
0

0
2
0
0
0

0
2
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
2
0
0
0

0
4
0
0
0

0
0
0
0
0

0
0
0
0
0

0
0
0
0
0

0
2
0
0
0

0
7
0
0
0

0
0
0
0
0

29
0
4
0
0

11
18
2
0
0

22
16
4
0
0

24
4
4
0
0

4
29
2
0
0

4
11
2
0
9

4
0
4
0
0

7
11
0
0
0

4
7
18
0
0

33
4
451
0
0

13
29
351
0
0

7
9
31
0
0

16
20
0
0
0

71
29
0
0
0

11
42
0
0
0

7
102
0
0
0

2
9
0
0

4
4
4
0
0

2
7
9
0
0

18
7
4
0
0

11
0
2
0
0

9
2
0
0
2

7
0
0
0
0

9
2
0
0
0

0
0
7
0

0
0
0
4
0

0
0
0
7
0

0
0
0
0
0

0
0
0
0
0

0
0
9
16
0

0
0
4
7
2

0
0
4
7
4

13
44
80
269
751

27
56
64
58
42

33
31
64
147
249

31
2
38
167
829

20
29
40
151
502

62
67
102
196
20

47
11
247
129
144

31
158
67
13
484

0
0
22

0
0
13

0
0
16

0
0
7

0
0
80

0
0
62

0
0
7

0
13
7

7
11
78
60
11000000000
15
Seto
0
0
0
0
0
0
2
2

0
36

0
13

0
11

0
18

4
211

4
24

2
0

7
18

0
0
0
0
0

0
0
0
0
2

0
0
0
0
7

0
0
0
0
0

0
0
0
2
7

0
0
0
27
13

0
0
0
0
76

0
0
0
0
20